Euphorbia thymifolia |
Euphorbia geyeri |
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gulf sandmat |
Geyer's sandmat, Geyer's spurge |
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Habit | Herbs, annual or perennial, with taproot. | Herbs, annual, with taproot. | ||||
Stems | prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose. |
prostrate or slightly ascending, loosely mat-forming, 4–25(–45) cm, glabrous. |
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Leaves | opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally. |
opposite; stipules usually distinct, occasionally connate basally on lower side, usually deeply parted into 3 or more filiform segments, 0.7–1.5 mm, glabrous; petiole 1–2 mm, glabrous; blade oblong to oblong-obovate or oblong-elliptic, 4–12 × 2–6 mm, base slightly asymmetric, angled or rounded, with one side usually expanded into small, rounded auricle, margins entire, apex usually truncate, occasionally emarginate, abaxial surface pale grayish green, both surfaces glabrous; only midvein conspicuous or venation obscurely pinnate (larger leaves). |
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Involucre | broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate. |
broadly campanulate, 1–1.5 × 0.7–0.9 mm, glabrous; glands 4, green to reddish, elliptic-oblong to nearly circular, slightly cupped to folded, 0.2–0.4 × 0.2–0.6 mm; appendages rudimentary to absent or white to reddish-tinged, usually rounded, sometimes pointed, (0–)0.5–1 × (0–)0.1–1.2 mm, distal margin entire or slightly toothed. |
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Staminate flowers | 3–5. |
5–20. |
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Pistillate flowers | ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2–0.6 mm, 2-fid nearly 1/2 length. |
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Capsules | conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm. |
globose-ovoid, 1.5–2 × 1.5–3 mm, glabrous; columella 1.5–1.9 mm. |
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Seeds | white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate. |
ashy white, ovoid, terete to bluntly subangled in cross section, 1.1–1.7 × 0.9–1.2 mm, smooth, with smooth brown line from top to bottom on adaxial side. |
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Cyathia | usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm. |
solitary or in small, cymose clusters at distal nodes; peduncle 1–2 mm. |
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Euphorbia thymifolia |
Euphorbia geyeri |
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Phenology | Flowering and fruiting midsummer–early fall. | |||||
Habitat | Disturbed areas, often near salt water. | |||||
Elevation | 0–20 m. (0–100 ft.) | |||||
Distribution |
FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] |
CO; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OK; SD; TX; UT; WI; WY; MB; n Mexico
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Discussion | Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). The two varieties of Euphorbia geyeri have been distinguished in large part by the presence of conspicuous involucral gland appendages in var. geyeri and the lack of appendages in var. wheeleriana. The two varieties are recognized here, but the variation in the size and presence of involucral gland appendages in the closely related E. bombensis suggests that this might be a somewhat variable character in this group of species. Euphorbia geyeri is widespread throughout the central United States in sandy soils. Populations at the eastern edge of the range are often considered adventive (for example, sandy soils along railroad grades in Michigan). Euphorbia geyeri resembles E. glyptosperma (both being entirely glabrous), but that species has serrulate leaves (near the apex) and strongly angled, transverse-ridged seeds whereas E. geyeri has entire leaves and smooth, rounded seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 291. | FNA vol. 12, p. 268. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
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Subordinate taxa | ||||||
Synonyms | Chamaesyce thymifolia | Chamaesyce geyeri | ||||
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Engelmann: Boston J. Nat. Hist. 5: 260. (1845) | ||||
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