FNA: Euphorbia thymifolia vs. Euphorbia curtisii

	Euphorbia thymifolia	Euphorbia curtisii
	gulf sandmat	Curtis' spurge, Sandhills spurge
Habit	Herbs, annual or perennial, with taproot.	Herbs, perennial, with spreading rootstock.
Stems	prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose.	erect or ascending, branched, solitary or few, previous year's dead stems not persistent, 20–40 cm, usually glabrous, rarely strigose to sericeous at nodes.
Leaves	opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally.	alternate; stipules to 0.1 mm; petiole to (0–)1–2 mm, glabrous or strigose to sericeous; blade usually linear, occasionally elliptic, rarely ovate, proximal often greatly reduced and often scalelike, 10–30 × 1.5–6 mm, base cuneate, margins entire, occasionally sparsely ciliate, apex rounded or broadly acute, abaxial surface glabrous or sparsely strigose to sericeous, adaxial surface glabrous; venation obscure, only midvein conspicuous.
Involucre	broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate.	campanulate, 1–1.2 × 1.3–1.5(–1.7) mm, glabrous or strigose to sericeous on distal 1/2; glands 5, green, reniform, 0.3 × 0.6 mm; appendages white, semicircular, 0.3–0.4 × 0.6–0.8 mm, entire.
Staminate flowers	3–5.	20–25.
Pistillate flowers	ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length.	ovary glabrous or sparsely strigose to sericeous; styles 0.6–1.1 mm, 2-fid at apex to 1/2 length.
Capsules	conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm.	globose, 2.5–3.2 × 4.3–5.1 mm, glabrous or sparsely strigose to sericeous; columella 2.4–3.1 mm.
Seeds	white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate.	usually gray to black, occasionally brown, ovoid-globose, 2.2 × 1.8 mm, smooth; caruncle absent.
Cyathia	usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm.	in terminal pleiochasia (fertile axillary branches occasionally present); peduncle 6.5–17 mm, filiform, glabrous.

	Euphorbia thymifolia	Euphorbia curtisii
Phenology	Flowering and fruiting midsummer–early fall.	Flowering and fruiting early spring–summer.
Habitat	Disturbed areas, often near salt water.	Xeric to dry oak or oak-pine scrub of sand hills, pine-oak woodlands, pine-oak savannas.
Elevation	0–20 m. (0–100 ft.)	0–200 m. (0–700 ft.)
Distribution	FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] [BONAP county map]	FL; GA; NC; SC [WildflowerSearch map] [BONAP county map]
Discussion	Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia curtisii is found in the Gulf and Atlantic coastal plains. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 291.	FNA vol. 12, p. 245.
Parent taxa	Euphorbiaceae > Euphorbia > sect. Anisophyllum	Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms	Chamaesyce thymifolia	E. eriogonoides, Tithymalopsis curtisii, T. eriogonoides
Name authority	Linnaeus: Sp. Pl. 1: 454. (1753)	Engelmann ex Chapman: Fl. South. U.S., 401. (1860)
Web links	Local Web sites: LA Plants WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia curtisii

gulf sandmat

Curtis' spurge, Sandhills spurge

Habit

Herbs, annual or perennial, with taproot.

Herbs, perennial, with spreading rootstock.

Stems

prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose.

erect or ascending, branched, solitary or few, previous year's dead stems not persistent, 20–40 cm, usually glabrous, rarely strigose to sericeous at nodes.

Leaves

opposite;

stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose;

petiole 0.5–1 mm, sparsely strigose-tomentulose;

blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally.

alternate;

stipules to 0.1 mm;

petiole to (0–)1–2 mm, glabrous or strigose to sericeous;

blade usually linear, occasionally elliptic, rarely ovate, proximal often greatly reduced and often scalelike, 10–30 × 1.5–6 mm, base cuneate, margins entire, occasionally sparsely ciliate, apex rounded or broadly acute, abaxial surface glabrous or sparsely strigose to sericeous, adaxial surface glabrous;

venation obscure, only midvein conspicuous.

Involucre

broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose;

glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm;

appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate.

campanulate, 1–1.2 × 1.3–1.5(–1.7) mm, glabrous or strigose to sericeous on distal 1/2;

glands 5, green, reniform, 0.3 × 0.6 mm;

appendages white, semicircular, 0.3–0.4 × 0.6–0.8 mm, entire.

Staminate flowers

3–5.

20–25.

Pistillate flowers

ovary densely strigose;

styles 0.4–0.5 mm, 2-fid 1/2 length.

ovary glabrous or sparsely strigose to sericeous;

styles 0.6–1.1 mm, 2-fid at apex to 1/2 length.

Capsules

conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose;

columella 0.6–1 mm.

globose, 2.5–3.2 × 4.3–5.1 mm, glabrous or sparsely strigose to sericeous;

columella 2.4–3.1 mm.

Seeds

white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate.

usually gray to black, occasionally brown, ovoid-globose, 2.2 × 1.8 mm, smooth;

caruncle absent.

Cyathia

usually in small, cymose clusters on congested, axillary branches;

peduncle (0–)0.1–0.3 mm.

in terminal pleiochasia (fertile axillary branches occasionally present);

peduncle 6.5–17 mm, filiform, glabrous.

Euphorbia thymifolia

Euphorbia curtisii

Phenology

Flowering and fruiting midsummer–early fall.

Flowering and fruiting early spring–summer.

Habitat

Disturbed areas, often near salt water.

Xeric to dry oak or oak-pine scrub of sand hills, pine-oak woodlands, pine-oak savannas.

Elevation

0–20 m. (0–100 ft.)

0–200 m. (0–700 ft.)

Distribution

FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia]

[BONAP county map]

FL; GA; NC; SC

[WildflowerSearch map]

[BONAP county map]

Discussion

Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia curtisii is found in the Gulf and Atlantic coastal plains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 291.

FNA vol. 12, p. 245.

Parent taxa

Euphorbiaceae > Euphorbia > sect. Anisophyllum

Euphorbiaceae > Euphorbia > sect. Alectoroctonum

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Synonyms

Chamaesyce thymifolia

E. eriogonoides, Tithymalopsis curtisii, T. eriogonoides

Name authority

Linnaeus: Sp. Pl. 1: 454. (1753)

Engelmann ex Chapman: Fl. South. U.S., 401. (1860)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

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Euphorbia thymifolia

Euphorbia curtisii

Euphorbia thymifolia

Euphorbia curtisii