Euphorbia thymifolia |
Euphorbia cordifolia |
|
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gulf sandmat |
heartleaf sandmat |
|
Habit | Herbs, annual or perennial, with taproot. | Herbs, annual, with taproot. |
Stems | prostrate, mat-forming, 15–30 cm, strigose to strigose-tomentulose. |
prostrate, occasionally mat-forming, 10–43 cm, glabrous. |
Leaves | opposite; stipules distinct or slightly connate at base, linear-subulate, entire or slightly parted, 0.9–1.2 mm, strigose-tomentulose; petiole 0.5–1 mm, sparsely strigose-tomentulose; blade broadly elliptic to narrowly oblong or ovate-lanceolate, 3–10 × 1.8–5 mm, base asymmetric, one side usually angled or rounded and the other truncate and expanded into small, rounded auricle, margins serrate (larger leaves) to serrulate (smaller leaves), apex blunt to acute, abaxial surface sparsely tomentulose to glabrate, adaxial surface glabrate; palmately veined at base, pinnate distally. |
opposite; stipules usually distinct, occasionally connate at base, filiform, 1–1.2(–2.8) mm, usually glabrous, rarely pilose; petiole 0.4–1 mm, usually glabrous; blade ovate to oblong, 4.4–12 × 2.6–7.6 mm, base asymmetric, cordate to rounded, margins entire, apex rounded to mucronulate, surfaces glabrous; only midvein conspicuous. |
Involucre | broadly obconic, becoming distended and distorted by base of partially included capsule, 0.4–0.8 × 0.3–0.5 mm, strigose; glands 4, red, slightly concave, ± unequal, subcircular to broadly oval, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, usually unequal, occasionally ± equal at distal nodes, elongated toward sinus, sometimes rudimentary, 0.1–0.4 × 0.1–0.3 mm, distal margin entire or crenulate. |
campanulate, 1–1.3 × 1–1.3 mm, glabrous; glands 4, yellowish to pink, elliptic, 0.3–0.5 × 0.5–1 mm; appendages whitish to pink, sometimes drying red, elliptic to ovoid, 1.1–1.5 × 1.2–1.9 mm, distal margin entire, retuse, or erose. |
Staminate flowers | 3–5. |
5–40. |
Pistillate flowers | ovary densely strigose; styles 0.4–0.5 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.5–0.8 mm, 2-fid nearly entire length. |
Capsules | conic to truncate-ovoid, scarcely exserted from involucre, base often remaining inside involucre and splitting one side of it during maturation, 0.9–1.2 × 1–1.2 mm, sparsely to moderately strigose; columella 0.6–1 mm. |
ovoid, 2–3 mm diam., glabrous; columella 1.2–2.7 mm. |
Seeds | white, tan underneath coat, ovoid to narrowly ovoid, sharply 4-angled in cross section, 0.8–0.9 × 0.4–0.6 mm, with 4 low transverse ridges often slightly extending into angles, not sulcate. |
gray or tan with dark brown mottling, ovoid, bluntly 3–4-angled in cross section, 1.8–2.1 × 1.2–1.4 mm, smooth to rugose. |
Cyathia | usually in small, cymose clusters on congested, axillary branches; peduncle (0–)0.1–0.3 mm. |
solitary at distal nodes; peduncle 0.9–3 mm. |
Euphorbia thymifolia |
Euphorbia cordifolia |
|
Phenology | Flowering and fruiting midsummer–early fall. | Flowering and fruiting summer–fall. |
Habitat | Disturbed areas, often near salt water. | Xeric oak-pine scrub, pine-barrens, sand barrens, sandy stream banks. |
Elevation | 0–20 m. (0–100 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
FL; LA; Mexico; South America; West Indies [Introduced in Asia, tropical Africa, Australia] |
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
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Discussion | Euphorbia thymifolia is a widespread tropical and subtropical weed. It is not certain where the species is native, but most likely it originated in the New World and then became widespread in the rest of the tropics. Euphorbia thymifolia is present in the flora area in southern Florida and coastal Louisiana, where it is likely adventive. Euphorbia thymifolia is generally similar to E. maculata but is characterized by its short pistillate pedicels and non-exserted capsules that remain largely enclosed by the involucre and by its unequal involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia cordifolia is easily identified by its cordate to rounded leaf base and distinctive filiform stipules. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 291. | FNA vol. 12, p. 265. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce thymifolia | Chamaesyce cordifolia |
Name authority | Linnaeus: Sp. Pl. 1: 454. (1753) | Elliott: Sketch. Bot. S. Carolina 2: 656. (1824) |
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