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Terlingua sandmat, Terlingua spurge

Roemer's spurge

Habit Herbs, annual, with slender taproot. Herbs, annual, with taproot.
Stems

prostrate to reclining, not mat-forming, 5–30 cm, glabrous.

erect, occasionally decumbent at base, branched, 15–30 cm, glabrous.

Leaves

opposite;

stipules usually distinct, occasionally connate basally on lower side of stem, subulate or scalelike, usually entire, occasionally 2-fid or margin sparsely ciliate, 0.4–1 mm, glabrous;

petiole 0.7–1.2(–1.5) mm, glabrous;

blade ovate, oblong, orbiculate, or obovate, 2–7.1 × 1–3.5 mm, base slightly asymmetric, rounded, margins entire, often revolute on drying, apex usually rounded, occasionally slightly emarginate, surfaces glabrous;

venation usually obscure, only midvein conspicuous.

petiole 2–5 mm;

blade oblanceolate to obovate, 5–20 × 5–10 mm (larger leaves in distal portion of stem), base cuneate to attenuate, margins entire, apex rounded, surfaces glabrous;

venation pinnate.

Involucre

usually turbinate-campanulate to hemispheric, occasionally suburceolate, 1–1.8 × 0.9–1.4 mm, glabrous;

glands 4, yellow-green to red-purple, sessile or short-stipitate, subcircular to slightly elliptic, 0.2–0.5 × 0.2–0.7 mm;

appendages absent or white to pink, semilunate or forming rim at edge of gland, (0–)0.1–0.4 × (0–)0.3–0.9 mm, entire or slightly crenate.

campanulate, 1.5–2 × 1–1.6 mm, glabrous;

glands 4, elliptic to trapezoidal, 0.4–0.6 × 0.7–1.2 mm;

horns slightly convergent, 0.1–0.3 mm.

Staminate flowers

15–36.

10–12.

Pistillate flowers

ovary glabrous;

styles 0.3–0.5 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.6–1 mm, 2-fid.

Capsules

broadly ovoid, 1.1–1.6 × 1.5–1.8 mm, glabrous;

columella 1.2–1.5 mm.

subglobose, 2–2.5 × 2.4–3 mm, slightly lobed;

cocci rounded, smooth, glabrous;

columella 1.5–1.8 mm.

Seeds

whitish, reddish brown beneath coat, ovate, 4-angled in cross section, 0.8–1.2 × 0.5–0.8 mm, with (2–)3(–5) deep transverse ridges.

brown, oblong-ovoid, 1.6–1.8 × 1.4–1.5 mm, with scattered, deep and broad pits;

caruncle conic, reniform, 0.3–0.4 × 0.4–0.6 mm.

Cyathia

usually solitary at distal nodes, rarely clustered on short, axillary branches;

peduncle 0.3–1.3 mm.

peduncle 0–0.9 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, 1–4 times 2-branched;

pleiochasial bracts ovate to oblong, similar in size to distal leaves;

dichasial bracts connate 1/2 length, reniform to semicircular, base truncate to perfoliate, margins entire, apex rounded to obtuse;

axillary cymose branches 0–3.

2n

= 28.

Euphorbia theriaca

Euphorbia roemeriana

Phenology Flowering and fruiting spring.
Habitat Rich calcareous soils, creek canyons.
Elevation 100–300 m. (300–1000 ft.)
Distribution
from FNA
NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. S. Correll and M. C. Johnston (1970) suggested that Euphorbia roemeriana, which is restricted to the eastern part of the Edwards Plateau, is the southern counterpart of E. commutata, to which it is morphologically very similar. However, molecular phylogenetic analyses indicate that it is most closely related to E. austrotexana and E. longicruris (J. A. Peirson et al. 2014). Euphorbia roemeriana can be distinguished from E. commutata and several of the other small, annual species of subg. Esula by its consistently connate dichasial bracts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral glands sessile, appendages absent.
var. theriaca
1. Involucral glands short-stipitate, appendages present, occasionally rudimentary.
var. spurca
Source FNA vol. 12, p. 290. FNA vol. 12, p. 308.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. theriaca var. spurca, E. theriaca var. theriaca
Synonyms Chamaesyce theriaca Tithymalus roemerianus
Name authority L. C. Wheeler: Rhodora 43: 242, plate 660, fig. A. (1941) Scheele: Linnaea 22: 151. (1849)
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