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Terlingua sandmat, Terlingua spurge

David's poinsettia, David's spurge, tooth poinsettia, tooth spurge

Habit Herbs, annual, with slender taproot. Herbs, annual, with taproot.
Stems

prostrate to reclining, not mat-forming, 5–30 cm, glabrous.

erect or ascending, 20–70 cm, both coarsely and sparsely hirsute and closely strigillose;

branches usually ± straight, occasionally proximal branches arcuate.

Leaves

opposite;

stipules usually distinct, occasionally connate basally on lower side of stem, subulate or scalelike, usually entire, occasionally 2-fid or margin sparsely ciliate, 0.4–1 mm, glabrous;

petiole 0.7–1.2(–1.5) mm, glabrous;

blade ovate, oblong, orbiculate, or obovate, 2–7.1 × 1–3.5 mm, base slightly asymmetric, rounded, margins entire, often revolute on drying, apex usually rounded, occasionally slightly emarginate, surfaces glabrous;

venation usually obscure, only midvein conspicuous.

usually opposite, occasionally alternate at distal nodes;

petiole 7–25 mm, strigose;

blade usually narrowly to broadly elliptic, occasionally lance-elliptic, 10–100 × 5–35 mm, base cuneate to attenuate, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface strigose with stiff, strongly tapered hairs, adaxial surface sparsely strigose-hirsute;

venation pinnate, midvein prominent.

Involucre

usually turbinate-campanulate to hemispheric, occasionally suburceolate, 1–1.8 × 0.9–1.4 mm, glabrous;

glands 4, yellow-green to red-purple, sessile or short-stipitate, subcircular to slightly elliptic, 0.2–0.5 × 0.2–0.7 mm;

appendages absent or white to pink, semilunate or forming rim at edge of gland, (0–)0.1–0.4 × (0–)0.3–0.9 mm, entire or slightly crenate.

cylindric, 2.5–3 × 1.3–1.8 mm, glabrous;

involucral lobes divided into 5–7 linear, papillate lobes;

gland 1, yellow-green, sessile and broadly attached, 0.9 × 1.3 mm, opening oblong, glabrous;

appendages absent.

Staminate flowers

15–36.

5–8.

Pistillate flowers

ovary glabrous;

styles 0.3–0.5 mm, 2-fid 1/2 length.

ovary glabrous or sparsely strigose;

styles 1 mm, 2-fid 1/2 to nearly entire length.

Capsules

broadly ovoid, 1.1–1.6 × 1.5–1.8 mm, glabrous;

columella 1.2–1.5 mm.

broadly ovoid, 2.9–3.3 × 4–4.8 mm, 3-lobed, glabrous;

columella 2.2–2.7 mm.

Seeds

whitish, reddish brown beneath coat, ovate, 4-angled in cross section, 0.8–1.2 × 0.5–0.8 mm, with (2–)3(–5) deep transverse ridges.

black to brown or pale gray, ovoid to triangular-ovoid, angular in cross section, 2.4–2.9 × 2.2–2.9 mm, low-tuberculate, tubercles irregularly arranged or in faint, transverse row;

caruncle 0.9–1.1 mm.

Cyathia

usually solitary at distal nodes, rarely clustered on short, axillary branches;

peduncle 0.3–1.3 mm.

peduncle 0.5–1 mm.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched;

pleiochasial bracts 2–4, often whorled, green with diffuse greenish white to mauve near base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts similar in shape to distal leaves but smaller, often highly reduced.

2n

= 28.

= 56.

Euphorbia theriaca

Euphorbia davidii

Phenology Flowering and fruiting summer–fall.
Habitat Forests, stream and riverbanks, prairies, roadsides and open disturbed areas.
Elevation 200–1500 m. (700–4900 ft.)
Distribution
from FNA
NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; AZ; CA; CO; FL; IA; ID; IL; IN; KS; KY; LA; MA; MI; MN; MO; NC; NE; NJ; NM; NY; OH; OK; SD; TN; TX; UT; VA; VT; WI; WV; WY; ON; QC; Mexico (Chihuahua, Coahuila, Sonora) [Introduced in South America, Eurasia (China, Russia), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia davidii is native from the southwestern United States and northern Mexico north through the southern Great Plains; it apparently is adventive elsewhere. The species is the weediest member of the E. dentata species group (following M. H. Mayfield 1997) and has become an agricultural weed in North America, South America (for example, Argentina), and in the Old World (particularly Australia and Russia). Euphorbia davidii can be distinguished from the closely similar E. dentata by its larger capsules and seeds, often more elliptic leaves, and shorter, stiffer hairs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Involucral glands sessile, appendages absent.
var. theriaca
1. Involucral glands short-stipitate, appendages present, occasionally rudimentary.
var. spurca
Source FNA vol. 12, p. 290. FNA vol. 12, p. 320.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Poinsettia
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. theriaca var. spurca, E. theriaca var. theriaca
Synonyms Chamaesyce theriaca
Name authority L. C. Wheeler: Rhodora 43: 242, plate 660, fig. A. (1941) Subils: Kurtziana 17: 125, figs. 1, 2H–J. (1984)
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