Euphorbia terracina |
Euphorbia misera |
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carnation spurge, Geraldton carnation weed, terracina spurge |
cliff spurge |
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Habit | Herbs, perennial or biennial, with taproot. | Shrubs, soft wooded, with woody rootstock. |
Stems | erect, unbranched or branched, 10–100 cm, glabrous. |
erect to ascending, often gnarled and scraggly, branched, with conspicuous knobby short shoots, 70–150 cm, puberulent-tomentose, bark grayish red to light gray. |
Leaves | petiole absent; blade linear, linear-lanceolate, oblong-elliptic, or obovate, 4–50 × 2–10 mm, base obtuse or truncate, margins finely serrulate, apex acute, obtuse, or truncate, sometimes mucronulate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
alternate, well spaced on long shoots or fasciculate on short shoots; stipules 0.6–1.1 mm; petiole 4–12(–19) mm, slender, puberulent to shortly pilose; blade oblong, ovate, orbiculate, elliptic, or obovate, 6–24 × 5–21 mm, base rounded to cuneate, margins entire, apex acute to obtuse, surfaces puberulent-tomentose; venation conspicuous. |
Involucre | cupulate to slightly turbinate, 1.1–2 × 1.3–1.5 mm, glabrous or puberulent; glands 4, elliptic to trapezoidal, 0.6–0.8 × 1–2 mm; horns slightly convergent to divergent, 1–2 mm. |
campanulate, 1.4–3.8 × 2.1–4.4 mm, puberulent-tomentose; glands 5, yellow to reddish, oblong to reniform, 0.7–1.3 × 1.1–2.6 mm; appendages green-yellow to yellowish or whitish, oblong to transversely oblong, 0.6–1.9 × 1.3–3.8 mm, crenulate to erose. |
Staminate flowers | 15–20. |
40–50. |
Pistillate flowers | ovary glabrous; styles 1–1.8 mm, 2-fid. |
ovary glabrous or puberulent; styles 1.6–2.7 mm, 2-fid at apex. |
Capsules | depressed-globose, 2.5–3 × 3–4.5 mm, deeply 3-lobed; cocci rounded to subangular, smooth, glabrous; columella 1.9–2.3 mm. |
oblate, 4.6–5.1 × 6.1–6.7 mm, usually glabrous or glabrescent, occasionally puberulent; columella 2.8–3.6 mm. |
Seeds | pale gray, subovoid, 1.6–2.4 × 1.3–1.8 mm, smooth; caruncle boat-shaped, 0.4–0.6 × 0.4–0.6 mm. |
grayish, subglobose to ovoid, rounded in cross section, 2.7–3.3 × 2.5–2.8 mm, foveolate; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 2–5, each 1–5 times 2-branched; pleiochasial bracts lanceolate, elliptic or ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to subreniform, base cuneate to cordate, margins finely serrulate, apex acute, obtuse, or rounded, sometimes mucronulate or cuspidate; axillary cymose branches 0–7. |
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Cyathia | peduncle 1–3 mm. |
usually solitary on short shoots, peduncle 1.8–10.5 mm, puberulent-tomentose. |
Euphorbia terracina |
Euphorbia misera |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round (but most prolific after winter rains). |
Habitat | Edges of cultivated fields and woodlands, roadsides, waste areas, pastures, coastal bluffs, dunes, riparian areas. | Rocky soils, sometimes in crevices of vertical cliff faces, coastal scrub, maritime desert scrub, arid desert scrub. |
Elevation | 0–300 m. (0–1000 ft.) | 0–400 m. (0–1300 ft.) |
Distribution |
CA; Europe [Introduced in North America; introduced also in Mexico, s Africa, Australia]
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CA; Mexico (Baja California, Baja California Sur, Sonora)
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Discussion | Euphorbia terracina is native to the Mediterranean region of Europe. This species is invasive and spreading rapidly, displacing native coastal scrub in southern California, and has been listed as a noxious weed by that state. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia misera is relatively infrequent within the flora area, known primarily from coastal southern California and the Channel Islands (although a relictual inland population occurs in the Little San Bernardino Mountains). The species has been considered worthy of conservation, but appears to be under little threat, especially in Mexico where it is frequent and often locally abundant. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 310. | FNA vol. 12, p. 249. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Tithymalus terracinus | |
Name authority | Linnaeus: Sp. Pl. ed. 2, 1: 654. (1762) | Bentham: Bot. Voy. Sulphur, 51. (1844) |
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