Euphorbia stictospora |
Euphorbia marginata |
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mat spurge, narrow-seed spurge, slimseed sandmat |
euphorbe marginée, mountain snow spurge, smoke-on-the-prairie, snow-on-the-mountain, variegated spurge, whitemargined spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | prostrate, often mat-forming, occasionally with ascending tips, 5–45 cm, densely and evenly pilose to lanate. |
erect, unbranched or branched, 30–85(–150) cm, pilose or glabrous. |
Leaves | opposite; stipules distinct or connate basally on one side of stem, entire or irregularly toothed or fringed, 0.5–1.2 mm, pilose to lanate; petiole 0.3–1.5 mm, pilose to lanate; blade usually oblong to oblong-obovate, occasionally nearly circular, 3–10(–15) × 2–5(–10) mm, base asymmetric, one side usually angled or rounded and other truncate-auriculate, margins minutely or conspicuously serrulate at least toward apex, apex usually broadly rounded to broadly acute, occasionally emarginate, abaxial surface often ± lighter green and without reddish spot, both surfaces sparsely to moderately pilose to lanate; 3-veined from base or venation obscure. |
alternate; stipules 0.1–0.3 mm; petiole 0.2–3 mm, glabrous or minutely pilose; blade broadly ovate to elliptic, 32–62(–82) × 18–28(–52) mm, base rounded to cuneate, margins entire, often white on distal leaves, apex acute, rarely mucronate, surfaces glabrous; venation obscure, only midvein conspicuous. |
Involucre | obconic, 0.7–1 × 0.4–0.6 mm, moderately to densely pilose to lanate; glands 4, reddish, ± unequal, oblong, 0.1 × 0.1–0.3 mm; appendages white to strongly pinkish or reddish tinged, often unequal, sometimes 1 to all absent, 0–0.3 × 0–0.4 mm, 3-lobed or rudimentarily 1-lobed, distal margin crenate. |
campanulate, 2.2–3.5 × 1.3–2.3 mm, margin between glands deeply divided into fringe of fimbriate lobes, pilose; glands 4–5, green to greenish yellow, reniform to subcircular, 0.7–1.1 × 1–1.6 mm; appendages white, orbiculate, 1.5–2.7 × 1.9–2.9(–3.6) mm, entire. |
Staminate flowers | 3–9. |
30–70. |
Pistillate flowers | ovary pilose to villous, hairs occasionally slightly appressed; styles 0.2–0.5 mm, unbranched or inconspicuously notched at tip. |
ovary pilose; styles 1–2.5 mm, 2-fid 1/2–2/3 length. |
Capsules | ovoid, 1.6–2.3 × 1.4–1.5 mm, moderately to densely villous with hairs usually slightly appressed, pubescence often concentrated on proximal 1/2 or along lobes; columella 1.5–2 mm. |
oblate, 3–5 × 3.5–7.5 mm, moderately to densely pilose; columella 3–4.1 mm. |
Seeds | light to dark brown, usually mottled, sometimes with thin, white coating, often wearing away irregularly, narrowly oblong-ovoid to ellipsoid, 3–4-angled in cross section, 1–1.5 × 0.5–0.6 mm, with short, irregularly interrupted furrows, appearing partially and irregularly few-ridged. |
orange-tan to gray, ovoid, 3.7–3.9 × 3–3.3 mm, rugose, with 2 transverse ridges (one dark orange to brown, other inconspicuous); caruncle absent. |
Cyathia | solitary at leaf nodes or in small, cymose clusters on congested, axillary branches; peduncle 0.7–2.5 mm. |
in terminal pleiochasia, dichasial bracts narrowly elliptic to oblanceolate with conspicuous white margins; peduncle 1.8–2.7(–22) mm, densely pilose. |
2n | = 56. |
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Euphorbia stictospora |
Euphorbia marginata |
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Phenology | Flowering and fruiting midsummer–early fall. | Flowering and fruiting summer–fall. |
Habitat | Open disturbed areas, rocky slopes. | Disturbed areas and grasslands. |
Elevation | 100–2100 m. (300–6900 ft.) | 0–1700 m. (0–5600 ft.) |
Distribution |
AZ; CO; IA; KS; MO; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua, Durango, San Luis Potosí)
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AR; CA; CO; CT; IA; IL; IN; KS; LA; MA; MD; MI; MN; MO; MT; NC; NE; NH; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; WI; WV; WY; MB; ON; QC; SK; c Mexico; s Mexico
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Discussion | Euphorbia stictospora has been recorded from New York, but this disjunct occurrence likely represents a waif or misidentification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia marginata is native to the central United States. The type specimen was collected by Meriwether Lewis along the Yellowstone River in southern Montana in 1806, and it has been reported to be native as far south as Arizona, New Mexico, and Texas, and as far east as southern Minnesota, western Iowa, and Missouri (G. Yatskievych 1999–2013, vol. 2). It is presumably naturalized outside of this area. Euphorbia marginata is widely cultivated as an ornamental for its showy, white-margined distal leaves, and it can escape locally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 289. | FNA vol. 12, p. 248. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Chamaesyce stictospora | E. bonplandii, Lepadena marginata |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) | Pursh: Fl. Amer. Sept. 2: 607. (1813) |
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