Euphorbia spathulata |
Euphorbia rosescens |
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prairie spurge, reticulate seed spurge, spatulate leaf spurge, warty spurge |
rosy-pink spurge, scrub spurge |
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Habit | Herbs, usually annual, rarely biennial, with taproot. | Herbs, perennial, with thickened rootstock. |
Stems | erect or ascending, unbranched or branched, 10–70 cm, glabrous. |
erect or ascending, 15–45 cm. |
Leaves | petiole absent or to 0.2 mm; blade oblanceolate, oblong-oblanceolate, spatulate, or cuneate, 10–50 × 6–11 mm, base broadly attenuate to rounded or shallowly cordate-clasping, margins finely serrulate (usually distally), apex usually rounded to obtuse, occasionally slightly retuse or obcordate proximally, bluntly mucronate, surfaces glabrous; venation pinnate, midvein prominent. |
petiole indistinct, blade narrowly elliptic, elliptic, narrowly oblong, or obovate, 25–55 × 5–21 mm, thick and fleshy, base attenuate, apex acute or mucronulate; only midvein evident. |
Involucre | campanulate to cupulate, 0.6–1(–1.5) × 0.8–1.2 mm, glabrous; glands 4–5, elliptic, oblong, to slightly reniform, 0.2–0.6 × 0.4–1 mm; horns absent. |
obconic to campanulate, 2.4–3.6 × 2.5–3.6 mm, lobes ovate to oblong, 0.7–1 mm, ciliate; glands green to yellow, oblong or trapezoidal, 0.9–1.7 × 1.9–2.5 mm, distal margins crenulate-erose. |
Staminate flowers | 3–10. |
25. |
Pistillate flowers | ovary glabrous; styles 0.8–1.5 mm, 2-fid. |
mature gynophore not seen, styles connate 1/2 length, 3.3–3.6 mm. |
Capsules | depressed-globose, 2–3.5 × 4 mm, 3-lobed; cocci rounded, verrucose, verrucae 0.1–0.2 mm, glabrous; columella 1.4–2.2 mm. |
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Seeds | red-brown to dark purple, occasionally ± glaucous, broadly ellipsoid-ovoid to nearly globose, 1.3–2.5 × 1.5–1.8 mm, smooth, reticulate, or finely low-ridged; caruncle irregularly reniform to round, subconic to lenticular, 0.3–0.4 × 0.5–0.6 mm. |
not seen. |
Cyathial | arrangement: terminal pleiochasial branches 3(–5), each 1–3 times 2-branched; pleiochasial bracts broadly ovate to ovate-oblong, shorter and wider than distal leaves; dichasial bracts distinct, broadly ovate, ovate-triangular, or ovate-elliptic, base cordate-clasping or subcordate to rounded, margins serrulate, apex rounded to obtuse or acute; axillary cymose branches (0–4)5–12. |
arrangement: terminal pleiochasial branches 3, 7–16 cm, 4–7 times 2-branched; pleiochasial bracts ovate to oblong, 17–31 × 13–14 mm, margins entire, apex acute, often also mucronulate; dichasial bracts ovate or lanceolate, 6–18 × 8–12 mm, margins entire, apex acute, often also mucronate; axillary cymose branches 1–2. |
Cyathia | peduncle 0.3–1(–1.5) mm. |
peduncle 3.9–6.8 mm. |
Mature | capsules not seen. |
|
Euphorbia spathulata |
Euphorbia rosescens |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–summer, rarely fall. |
Habitat | Forests, fallow fields, prairies, pastures, glades, stream banks, waste places, roadsides. | Xeric oak and pine scrub, mostly on white sands, disturbed habitats. |
Elevation | 0–3500 m. (0–11500 ft.) | 20–50 m. (100–200 ft.) |
Distribution |
AL; AR; AZ; CA; CO; DC; FL; GA; IA; ID; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; MT; NC; ND; NE; NM; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; ON; Mexico (Chihuahua, Sonora); s South America
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FL |
Discussion | As treated here, Euphorbia spathulata is a wide-ranging and variable species. J. B. S. Norton (1900) recognized a number of segregates (for example, E. arkansana, E. dictyosperma, and E. obtusata) all of which are included here in a broadly defined E. spathulata. The only segregate species that has been widely recognized in regional floras is E. obtusata (for example, M. L. Fernald 1950; T. S. Cooperrider 1995; G. Yatskievych 1999–2013, vol. 2). Authors have generally distinguished the eastern North American E. obtusata from the western E. spathulata by the former's larger seeds (1.7–2.3 mm versus 1.5–1.7 mm) with smooth (versus reticulate) surfaces, larger involucres, red (versus yellow) involucral glands, and cordate-clasping (versus rounded to subcordate) dichasial bracts. Examination of specimens of E. spathulata in the broad sense from throughout North America showed that there is some geographic patterning to seed size and surface sculpturing, but the variation does not segregate cleanly into two discrete taxa. Plants from western North America typically have small seeds (1.5–1.7 mm) with reticulate surfaces, although some western individuals have seeds 1.8–1.9 mm long with reticulate surfaces. Plants from Texas generally have small seeds (1.5–1.6 mm) but with the surfaces either reticulate or completely smooth. Plants from adjacent Louisiana have small seeds with faintly reticulate to almost bumpy surfaces. Plants from eastern North America have larger seeds (2–2.3 mm) with usually smooth surfaces, although individuals from Tennessee and the Carolinas have faintly reticulate surfaces. Involucre height, gland color, and the shape of the dichasial bracts do not segregate with seed size as previous treatments have suggested. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia rosescens is a narrow-endemic, gap-specialist known only from the southern portion of the Lake Wales Ridge in Highlands County. Based on leaf characteristics, it appears to be most similar to E. telephioides. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 309. | FNA vol. 12, p. 316. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Nummulariopsis |
Sibling taxa | ||
Synonyms | E. arkansana, E. dictyosperma, E. obtusata, Galarhoeus arkansanus, G. obtusatus, Tithymalus arkansanus, T. dictyospermus, T. obtusatus, T. spathulatus | |
Name authority | Lamarck: in J. Lamarck et al., Encycl. 2: 428. (1788) | E. L. Bridges & Orzell: Lundellia 5: 71, fig. 4. (2002) |
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