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fringe spurge, shaggy spurge, Yuma sandmat

perennial sandmat, Terlingua spurge

Habit Herbs, annual, with slender taproot. Herbs, perennial, with strongly thickened, woody rootstock.
Stems

prostrate, mat-forming, 5–50 cm, villous with glistening glandular hairs.

erect, 7–45 cm, glabrous.

Leaves

opposite;

stipules distinct, filiform, rudimentary to 0.2 mm, glabrous or sparsely villous with glistening glandular hairs;

petiole 0.5–1.5 mm, villous;

blade oblong, ovate, or elliptic, 3–7 × 2–4 mm, base asymmetric, rounded, margins entire, apex obtuse, surfaces villous; weakly 3-veined from base, commonly only midvein conspicuous.

opposite;

stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous;

petiole 0.8–2 mm, glabrous;

blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous.

Involucre

campanulate or urceolate, 0.7–1 × 0.5–0.8 mm, villous;

glands 4, red to pink, oblong to slightly reniform, 0.1–0.2 × 0.2–0.3 mm;

appendages white to pink, deeply incised into 3–6 triangular to subulate, attenuate, acute segments, 0.3–0.6 × 0.6–1 mm, segments entire.

broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous;

glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm;

appendages absent.

Staminate flowers

3–7.

35–45.

Pistillate flowers

ovary villous;

styles 0.3–0.4 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.7–0.9 mm, 2-fid nearly entire length.

Capsules

subglobose to ovoid, 1–1.2 mm diam., villous;

columella 0.9–1.1 mm.

subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous;

columella 2.2–2.7 mm.

Seeds

pink to light gray, narrowly ovoid, 4-angled in cross section, 0.8–1 × 0.5–0.6 mm, dimpled or with faint transverse ridges that do not pass through abaxial keel.

white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled.

Cyathia

solitary at distal nodes, nodes often congested toward tips of branches;

peduncle 0.2–1.6 mm.

solitary at distal nodes or at nodes of short, axillary branches;

peduncle 1.8–3 mm.

Euphorbia setiloba

Euphorbia perennans

Phenology Flowering nearly year-round in response to sufficient moisture. Flowering and fruiting spring–fall.
Habitat Desert scrub, blackbrush scrub, Joshua tree woodlands, grasslands, often in sandy areas. Desert scrub, on cretaceous and gypseous clay, limestone hills and flats.
Elevation 20–1600 m. (100–5200 ft.) 900–1200 m. (3000–3900 ft.)
Distribution
from FNA
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Durango, Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua)
[BONAP county map]
Discussion

Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011).

Euphorbia perennans is known in the flora area only from Brewster County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 288. FNA vol. 12, p. 282.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce setiloba Chamaesyce perennans
Name authority Engelmann: in War Department [U.S.], Pacif. Railr. Rep. 5(2): 364. (1857) (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960)
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