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fringe spurge, shaggy spurge, Yuma sandmat

Florida pineland spurge

Habit Herbs, annual, with slender taproot. Herbs, perennial, with thickened rootstock.
Stems

prostrate, mat-forming, 5–50 cm, villous with glistening glandular hairs.

erect or ascending, 15–40 cm.

Leaves

opposite;

stipules distinct, filiform, rudimentary to 0.2 mm, glabrous or sparsely villous with glistening glandular hairs;

petiole 0.5–1.5 mm, villous;

blade oblong, ovate, or elliptic, 3–7 × 2–4 mm, base asymmetric, rounded, margins entire, apex obtuse, surfaces villous; weakly 3-veined from base, commonly only midvein conspicuous.

petiole absent or indistinct, blade linear to linear-elliptic or lanceolate, (25–)30–60(–115) × 1.5–14(–15) mm, chartaceous, base attenuate, apex acuminate, acute, or rounded and mucronate;

only midvein evident.

Involucre

campanulate or urceolate, 0.7–1 × 0.5–0.8 mm, villous;

glands 4, red to pink, oblong to slightly reniform, 0.1–0.2 × 0.2–0.3 mm;

appendages white to pink, deeply incised into 3–6 triangular to subulate, attenuate, acute segments, 0.3–0.6 × 0.6–1 mm, segments entire.

obconic or campanulate, 1.3–3.2 × 1.6–3.6 mm, lobes ovate, oblong, or nearly rectangular, 0.6–1 mm, laciniate-ciliate;

glands red to greenish, oblong or nearly circular, 0.8–1 × 1.1–2 mm, distal margins crenulate-erose, undulate, or entire.

Staminate flowers

3–7.

20–25.

Pistillate flowers

ovary villous;

styles 0.3–0.4 mm, 2-fid 1/2 length.

gynophore exserted 2.6–6.1 mm, calyxlike lobes triangular to subulate, 0.6–1.4 mm;

styles connate 1/8 length, 1.3–2.2 mm.

Capsules

subglobose to ovoid, 1–1.2 mm diam., villous;

columella 0.9–1.1 mm.

oblate-ovoid, 5.1–6.1 × 6.9–8.6 mm, 3-lobed;

columella 3.9–4.7 mm.

Seeds

pink to light gray, narrowly ovoid, 4-angled in cross section, 0.8–1 × 0.5–0.6 mm, dimpled or with faint transverse ridges that do not pass through abaxial keel.

brown to blackish, ovoid-globose, circular or faintly 3- or 4-angled in cross section, 2.9–3.4 × 2.7–3 mm, smooth, base flattened or rounded, apex rounded or with inconspicuous blunt point.

Cyathia

solitary at distal nodes, nodes often congested toward tips of branches;

peduncle 0.2–1.6 mm.

peduncle 6–24 mm (often exceeding dichasial bracts).

Cyathial

arrangement: terminal pleiochasial branches 2–3, 5–18 cm, 3–8 times 2-branched;

pleiochasial bracts linear-lanceolate or narrowly ovate, 25–49 × 5–9 mm, margins entire, apex acute or acuminate;

dichasial bracts ovate or lanceolate, 6–21 × 2–6 mm, margins entire, apex acute or acuminate;

axillary cymose branches 1–5.

Euphorbia setiloba

Euphorbia inundata

Phenology Flowering nearly year-round in response to sufficient moisture.
Habitat Desert scrub, blackbrush scrub, Joshua tree woodlands, grasslands, often in sandy areas.
Elevation 20–1600 m. (100–5200 ft.)
Distribution
from FNA
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Durango, Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; FL; GA; MS
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Euphorbia inundata has a disjunct distribution in the southeastern United States. Allopatric, narrow-leaved populations from the west-central Florida peninsula are segregated as var. garrettii (E. L. Bridges and S. L. Orzell 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades usually linear to narrowly elliptic or narrowly lanceolate, rarely oblanceolate, (3–)4–14(–15) mm wide, length usually 5–10(–25) times width; apex obtuse to short-acute; se Mississippi to ne Florida.
var. inundata
1. Leaf blades narrowly linear, linear, or narrowly linear-lanceolate, 1.5–3.5(–4.5) mm wide, length 15–20(–50) times width, apex short-acute to acuminate; wc Florida.
var. garrettii
Source FNA vol. 12, p. 288. FNA vol. 12, p. 314.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Nummulariopsis
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. inundata var. garrettii, E. inundata var. inundata
Synonyms Chamaesyce setiloba Galarhoeus inundatus
Name authority Engelmann: in War Department [U.S.], Pacif. Railr. Rep. 5(2): 364. (1857) Torrey ex Chapman: Fl. South. U.S., 402. (1860)
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