Euphorbia serpens |
Euphorbia schizoloba |
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creeping spurge, Matted sandmat, Matted spurge, round-leafed spurge |
Mojave spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, perennial, with thick rootstock. |
Stems | prostrate, frequently mat-forming and rooting at nodes, 15–50 cm, glabrous. |
slender, ascending, often sinuous, many, densely branched near base, 10–50 cm, usually glabrous, usually glaucous. |
Leaves | opposite; stipules connate into conspicuous, deltate scale, white to pink, membranaceous, 0.5–1(–1.2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 2–7(–9) × 2–6 mm, base asymmetric, rounded to subcordate, margins entire, apex rounded, surfaces without red blotch, glabrous; usually only midvein conspicuous. |
petiole 0–1 mm; blade broadly oblanceolate to obovate, 10–20 × 3–9 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate to cuspidate, surfaces usually glabrous, usually glaucous; venation pinnate, sometimes obscure, midvein prominent. |
Involucre | campanulate to turbinate, 0.3–0.7(–1) × 0.4–0.6 mm, glabrous; glands 4, yellow, oblong, 0.1 × 0.2 mm; appendages white to pinkish, equal, forming narrow rim at edge of gland, 0.1–0.2 × 0.2–0.3 mm, distal margin entire or crenulate. |
campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, glabrous; glands 4, irregularly semicircular to trapezoidal or elliptic-truncate, 0.8–1.5 × 1–2.2, margins strongly crenate or dentate; horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin. |
Staminate flowers | 5–10. |
12–20. |
Pistillate flowers | ovary glabrous; styles 0.2 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 1–1.2 mm, 2-fid. |
Capsules | broadly ovoid, oblate, or subglobose, 1.3–1.4 × 1.3–1.7 mm, glabrous; columella (0.9–)1–1.2 mm. |
oblong-ovoid, 3.5–4 × 3.8–5 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.3–3.8 mm. |
Seeds | white to gray or light pink, ovoid, bluntly 3–4-angled in cross section, 0.7–1.1 × 0.4–0.7 mm, smooth. |
gray to whitish, oblong cylindric, 2–3 × 1.5 mm, irregularly shallowly pitted to almost smooth; caruncle conic, 0.6 × 0.6 mm. |
Cyathia | solitary at distal nodes; peduncle 0.5–1(–2.5) mm. |
peduncle 0.3–1 mm. |
Cyathial | arrangement: terminal pleiochasial branches (1–)3–4(–5), each 1–3 times 2-branched; pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves; dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate; axillary cymose branches 0–2(–4). |
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Euphorbia serpens |
Euphorbia schizoloba |
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Phenology | Flowering and fruiting year-round (in warmer areas) or summer (in temperate regions). | Flowering and fruiting spring–summer. |
Habitat | Mostly sandy or well-drained soils, desert scrub, coastal scrub, chaparral, oak and juniper woodlands, sand dunes, riparian forests, mesquite grasslands, prairies, coniferous and deciduous hardwood forests, disturbed areas. | Desert mountains and canyon slopes, rocky and gravelly soils. |
Elevation | 0–2000 m. (0–6600 ft.) | 500–1800 m. (1600–5900 ft.) |
Distribution |
AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; MT; ND; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; VT; WY; ON; SK; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Australia]
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AZ; CA; NV
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Discussion | Euphorbia serpens is one of the most widespread species of the genus in the New World. While it may be indigenous to a portion of the flora area, probably in the warmer, southern part of its range, it is weedy and has likely been introduced in many parts of the flora area, such as Canada and the eastern United States. It is also widely distributed in the Old World, where it is certainly introduced. The strictly prostrate habit with stems rooting at the nodes is characteristic. Euphorbia serpens is often confused with E. albomarginata, a species distributed in the southwestern United States; in addition to the features mentioned in the key, an easy and reliable way to distinguish between them is by the size of the involucral gland appendages: those of E. albomarginata are conspicuous to the naked eye, whereas those of E. serpens are inconspicuous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia schizoloba is a desert perennial that occurs on bluffs and ledges in the Mojave and Sonoran Deserts. Sparsely pubescent plants of E. schizoloba are known from Arizona and are best represented by several collections from the Mazatzal and Sierra Ancha mountains in Gila County from between 1000 to 1800 meters. George Engelmann published two names for this species almost simultaneously in 1861 (E. schizoloba and E. incisa). Although the authors have not been able to determine which publication has priority, Engelmann himself cited E. incisa as a synonym of E. schizoloba (in W. H. Brewer et al. 1876–1880, vol. 2), as did J. B. S. Norton (1899). Also, the type specimen at MO was annotated by Engelmann as E. schizoloba, and there is no mention of the name E. incisa on the sheet. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 286. | FNA vol. 12, p. 309. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce serpens | E. incisa, Tithymalus incisus, T. schizolobus |
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 41; 2(qto.): 52. (1817) | Engelmann: Proc. Amer. Acad. Arts 5: 173. (1861) |
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