Euphorbia serpens |
Euphorbia longicruris |
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creeping spurge, Matted sandmat, Matted spurge, round-leafed spurge |
wedge-leaf spurge |
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Habit | Herbs, annual, with slender taproot. | Herbs, annual, with taproot. |
Stems | prostrate, frequently mat-forming and rooting at nodes, 15–50 cm, glabrous. |
erect, usually unbranched, occasionally branched later in season, 5–25 cm, glabrous. |
Leaves | opposite; stipules connate into conspicuous, deltate scale, white to pink, membranaceous, 0.5–1(–1.2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 2–7(–9) × 2–6 mm, base asymmetric, rounded to subcordate, margins entire, apex rounded, surfaces without red blotch, glabrous; usually only midvein conspicuous. |
petiole 0–0.5 mm; blade cuneate-spatulate to obovate, 5–15 × 2–6 mm, base broadly attenuate, margins entire, apex rounded to obtuse, mucronate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | campanulate to turbinate, 0.3–0.7(–1) × 0.4–0.6 mm, glabrous; glands 4, yellow, oblong, 0.1 × 0.2 mm; appendages white to pinkish, equal, forming narrow rim at edge of gland, 0.1–0.2 × 0.2–0.3 mm, distal margin entire or crenulate. |
campanulate, 1.5–2 × 1–1.5 mm, glabrous; glands 4, crescent-shaped to elliptic, 0.4–0.8 × 0.8–1.1 mm; horns divergent, 0.5–0.8 mm. |
Staminate flowers | 5–10. |
10–15. |
Pistillate flowers | ovary glabrous; styles 0.2 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.5–0.6 mm, 2-fid. |
Capsules | broadly ovoid, oblate, or subglobose, 1.3–1.4 × 1.3–1.7 mm, glabrous; columella (0.9–)1–1.2 mm. |
ovoid-globose, 2–2.8 × 2.5–3 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 1.6–2.1 mm. |
Seeds | white to gray or light pink, ovoid, bluntly 3–4-angled in cross section, 0.7–1.1 × 0.4–0.7 mm, smooth. |
gray to purple-gray or sometimes nearly black, oblong, 1.3–1.6 × 0.9–1.2 mm, strongly small-pitted; caruncle umbonate, depressed-conic, 0.5 × 0.7 mm. |
Cyathia | solitary at distal nodes; peduncle 0.5–1(–2.5) mm. |
peduncle 0.3–0.5 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each many times 2-branched; pleiochasial bracts obovate, similar in size to distal leaves; dichasial bracts basally subconnate, strongly imbricate and often obscuring internodes, reniform to semiorbiculate, base cordate, margins entire, apex rounded; axillary cymose branches 0–5. |
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Euphorbia serpens |
Euphorbia longicruris |
|
Phenology | Flowering and fruiting year-round (in warmer areas) or summer (in temperate regions). | Flowering and fruiting spring. |
Habitat | Mostly sandy or well-drained soils, desert scrub, coastal scrub, chaparral, oak and juniper woodlands, sand dunes, riparian forests, mesquite grasslands, prairies, coniferous and deciduous hardwood forests, disturbed areas. | Grasslands, open prairies, sites with rocky, usually calcareous soils. |
Elevation | 0–2000 m. (0–6600 ft.) | 300–800 m. (1000–2600 ft.) |
Distribution |
AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; MT; ND; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; VT; WY; ON; SK; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Australia]
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AR; KS; OK; TX |
Discussion | Euphorbia serpens is one of the most widespread species of the genus in the New World. While it may be indigenous to a portion of the flora area, probably in the warmer, southern part of its range, it is weedy and has likely been introduced in many parts of the flora area, such as Canada and the eastern United States. It is also widely distributed in the Old World, where it is certainly introduced. The strictly prostrate habit with stems rooting at the nodes is characteristic. Euphorbia serpens is often confused with E. albomarginata, a species distributed in the southwestern United States; in addition to the features mentioned in the key, an easy and reliable way to distinguish between them is by the size of the involucral gland appendages: those of E. albomarginata are conspicuous to the naked eye, whereas those of E. serpens are inconspicuous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia longicruris is quite similar to the other small, annual members of subg. Esula in the south-central United States and can best be distinguished from those species by its imbricate dichasial bracts that form little tufts of overlapping leaves at the ends of the pleiochasial branches. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 286. | FNA vol. 12, p. 304. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce serpens | Tithymalus longicruris |
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 41; 2(qto.): 52. (1817) | Scheele: Linnaea 22: 152. (1849) |
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