Euphorbia serpens |
Euphorbia fendleri |
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creeping spurge, Matted sandmat, Matted spurge, round-leafed spurge |
Fendler's sandmat |
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Habit | Herbs, annual, with slender taproot. | Herbs, perennial, with woody, thickened taproot. |
Stems | prostrate, frequently mat-forming and rooting at nodes, 15–50 cm, glabrous. |
usually prostrate, decumbent, or ascending, very rarely erect, often densely clustered from top of woody crown, 5–12 cm, glabrous. |
Leaves | opposite; stipules connate into conspicuous, deltate scale, white to pink, membranaceous, 0.5–1(–1.2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 2–7(–9) × 2–6 mm, base asymmetric, rounded to subcordate, margins entire, apex rounded, surfaces without red blotch, glabrous; usually only midvein conspicuous. |
usually opposite, rarely whorled; stipules distinct, narrowly linear, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade usually orbiculate to ovate, rarely almost lanceolate, 3–8 × 2.5–7 mm, base slightly asymmetric, slightly cordate to rounded or obtuse, margins entire, apex rounded to obtuse, surfaces glabrous; obscurely 3–5-veined from base, only midvein conspicuous. |
Involucre | campanulate to turbinate, 0.3–0.7(–1) × 0.4–0.6 mm, glabrous; glands 4, yellow, oblong, 0.1 × 0.2 mm; appendages white to pinkish, equal, forming narrow rim at edge of gland, 0.1–0.2 × 0.2–0.3 mm, distal margin entire or crenulate. |
campanulate to turbinate or broadly cupulate, 1.1–1.7 × 1.2–1.8 mm, glabrous; glands 4, yellow-green to reddish, elliptic to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages absent or white, rarely pink, often unequal, lunate to flabellate or sometimes forming crenate margin along gland, (0–)0.1–0.6 × (0–)0.5–1.5 mm, distal margin entire or toothed. |
Staminate flowers | 5–10. |
25–35. |
Pistillate flowers | ovary glabrous; styles 0.2 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
Capsules | broadly ovoid, oblate, or subglobose, 1.3–1.4 × 1.3–1.7 mm, glabrous; columella (0.9–)1–1.2 mm. |
depressed-globose, 2–2.4 × 2.2–2.5 mm, glabrous; columella 1.7–2.1 mm. |
Seeds | white to gray or light pink, ovoid, bluntly 3–4-angled in cross section, 0.7–1.1 × 0.4–0.7 mm, smooth. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.7–2 × 1–1.2 mm, smooth to slightly wrinkled. |
Cyathia | solitary at distal nodes; peduncle 0.5–1(–2.5) mm. |
solitary at distal nodes; peduncle 0.7–1.2 mm. |
2n | = 28. |
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Euphorbia serpens |
Euphorbia fendleri |
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Phenology | Flowering and fruiting year-round (in warmer areas) or summer (in temperate regions). | Flowering and fruiting spring–fall. |
Habitat | Mostly sandy or well-drained soils, desert scrub, coastal scrub, chaparral, oak and juniper woodlands, sand dunes, riparian forests, mesquite grasslands, prairies, coniferous and deciduous hardwood forests, disturbed areas. | Mountain slopes, desert scrub, pinyon-juniper woodlands, hills, canyons, grasslands, washes, roadsides, dry crevices in limestone, often in gravel and sand. |
Elevation | 0–2000 m. (0–6600 ft.) | 500–2600 m. (1600–8500 ft.) |
Distribution |
AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; MT; ND; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; VT; WY; ON; SK; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Australia]
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AZ; CA; CO; KS; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Coahuila)
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Discussion | Euphorbia serpens is one of the most widespread species of the genus in the New World. While it may be indigenous to a portion of the flora area, probably in the warmer, southern part of its range, it is weedy and has likely been introduced in many parts of the flora area, such as Canada and the eastern United States. It is also widely distributed in the Old World, where it is certainly introduced. The strictly prostrate habit with stems rooting at the nodes is characteristic. Euphorbia serpens is often confused with E. albomarginata, a species distributed in the southwestern United States; in addition to the features mentioned in the key, an easy and reliable way to distinguish between them is by the size of the involucral gland appendages: those of E. albomarginata are conspicuous to the naked eye, whereas those of E. serpens are inconspicuous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia fendleri is similar to E. chaetocalyx and may sometimes be confused with that species. Its prostrate to decumbent or ascending stems and small, ovate to orbiculate leaves distinguish it from E. chaetocalyx. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. fendleri from E. chaetocalyx, but those characters appear highly variable and of little taxonomic utility; somewhat intermediate individuals occur in western Arizona, New Mexico, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 286. | FNA vol. 12, p. 267. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce serpens | Chamaesyce fendleri |
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 41; 2(qto.): 52. (1817) | Torrey & A. Gray: in War Department [U.S.], Pacif. Railr. Rep. 2(4): 175. (1857) |
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