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sun spurge

woodland spurge

Habit Herbs, perennial, with moniliform tuberous rootstock. Herbs, perennial, with thick rootstock.
Stems

erect, 5–20(–30) cm, usually glabrous, occasionally puberulent;

branches ± straight.

erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent.

Leaves

alternate;

petiole 0–2 mm, glabrous or strigose;

blade linear-lanceolate to ovate or broadly elliptic, 25–50 × 3–20 mm, unlobed, base rounded (tapered to petiole), margins with few glandular teeth, strigillose, flat to revolute, apex acute, abaxial surface coarsely strigose, adaxial surface strigose-hirsute;

venation pinnate, midvein prominent.

petiole 0–1 mm;

blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous;

venation inconspicuous, only midvein prominent.

Involucre

broadly globose-cupulate, 1.7–2.1 × 2.2–2.5 mm, glabrous or puberulent;

involucral lobes divided into triangular segments;

glands 1–4(–5), white, sessile and broadly attached, 1.1 × 1.4 mm, opening oblong, glabrous;

appendages absent.

cupulate, 2–2.2 × 1.3–1.8 mm, glabrous;

glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate;

horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin.

Staminate flowers

20–25.

10–20.

Pistillate flowers

ovary glabrous or puberulent, styles 3–4 mm, 2-fid 1/2 to nearly entire length.

ovary glabrous or puberulent;

styles 0.7–1 mm, 2-fid.

Capsules

depressed-globose, 3.8–5 × 4–5 mm, 3-lobed, glabrous or puberulent;

columella 3.6–4.5 mm.

ovoid, 3.5–4 × 4–4.5 mm, 3-lobed;

cocci rounded, smooth to slightly rugose, glabrous;

columella 3.2–3.5 mm.

Seeds

white, mottled brown to gray, ellipsoid, rounded in cross section, 4–4.6 × 2.4–3.2 mm, smoothly and broadly pitted or grooved;

caruncle 0.1 mm.

gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted;

caruncle conic, 0.6 × 0.7 mm.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched (often highly condensed);

pleiochasial bracts 6–8(–10), as tight involucrate whorl, wholly white to pale pink or red, usually narrower than distal leaves;

dichasial bracts linear and highly reduced.

arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched;

pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves;

dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate;

axillary cymose branches 0–4.

Cyathia

peduncle 2–5.5 mm.

peduncle 0.3–0.9 mm.

Euphorbia radians

Euphorbia lurida

Phenology Flowering and fruiting spring–summer. Flowering and fruiting spring–summer.
Habitat Pinyon-juniper woodlands, oak savannas, desert grasslands and scrub. Open pine-oak forests, dry slopes and canyons.
Elevation 700–2500 m. (2300–8200 ft.) 1300–2800 m. (4300–9200 ft.)
Distribution
from FNA
AZ; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; NV; UT; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia radians is widely distributed but scattered from the Sonoran and Chihuahuan deserts south to Oaxaca in Mexico. The species is distinct among species in sect. Poinsettia in the flora area in its precocious habit, often flowering before the leaves emerge.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 324. FNA vol. 12, p. 305.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Poinsettia radians E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens
Name authority Bentham: Pl. Hartw., 8. (1839) Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861)
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