Euphorbia radians |
Euphorbia humistrata |
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sun spurge |
spreading sandmat |
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Habit | Herbs, perennial, with moniliform tuberous rootstock. | Herbs, annual, with taproot. |
Stems | erect, 5–20(–30) cm, usually glabrous, occasionally puberulent; branches ± straight. |
prostrate to ascending, usually mat-forming and rooting at nodes, 5–45 cm, sparsely to moderately villous to pilose (densely on young growth). |
Leaves | alternate; petiole 0–2 mm, glabrous or strigose; blade linear-lanceolate to ovate or broadly elliptic, 25–50 × 3–20 mm, unlobed, base rounded (tapered to petiole), margins with few glandular teeth, strigillose, flat to revolute, apex acute, abaxial surface coarsely strigose, adaxial surface strigose-hirsute; venation pinnate, midvein prominent. |
opposite; stipules distinct, linear-subulate, often irregularly 2- or 3-lobed, 1–1.3 mm, sparsely villous to pilose; petiole 0.5–1.5 mm, sparsely to moderately villous to pilose; blade oblong-ovate to ovate-elliptic or oblong-elliptic, 4–18 × 2.5–8 mm, base strongly asymmetric, one side angled and other rounded to auriculate, margin on longer side serrulate, on shorter side subentire, apex rounded or broadly acute, abaxial surface pale grayish green, sparsely lanulose, adaxial surface usually with irregular reddish streak along midvein, usually glabrate, rarely sparsely lanulose; palmately veined at base, pinnate distally. |
Involucre | broadly globose-cupulate, 1.7–2.1 × 2.2–2.5 mm, glabrous or puberulent; involucral lobes divided into triangular segments; glands 1–4(–5), white, sessile and broadly attached, 1.1 × 1.4 mm, opening oblong, glabrous; appendages absent. |
obconic, 0.8–1 × 0.6–0.8 mm, sparsely villous to pilose; glands 4, green to yellow-green (turning pink with age), usually ± unequal, narrowly oblong, 0.1–0.2 × 0.2–0.5 mm; appendages white to reddish tinged, lunate, ± irregular and variable in shape, 0.1–0.3 × 0.2–1.5 mm, distal margin crenulate. |
Staminate flowers | 20–25. |
2–5. |
Pistillate flowers | ovary glabrous or puberulent, styles 3–4 mm, 2-fid 1/2 to nearly entire length. |
ovary short-sericeous; styles 0.5–0.8 mm, 2-fid 1/2 length. |
Capsules | depressed-globose, 3.8–5 × 4–5 mm, 3-lobed, glabrous or puberulent; columella 3.6–4.5 mm. |
ovoid, well exserted from involucre at maturity, 1.3–1.5 × 1.2–1.6 mm, sparsely to moderately short-sericeous; columella 0.9–1.2 mm. |
Seeds | white, mottled brown to gray, ellipsoid, rounded in cross section, 4–4.6 × 2.4–3.2 mm, smoothly and broadly pitted or grooved; caruncle 0.1 mm. |
white to light brown, oblong-ovoid, bluntly angular in cross section, 0.8–1.2 × 0.5–0.9 mm, smooth or papillate. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched (often highly condensed); pleiochasial bracts 6–8(–10), as tight involucrate whorl, wholly white to pale pink or red, usually narrower than distal leaves; dichasial bracts linear and highly reduced. |
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Cyathia | peduncle 2–5.5 mm. |
solitary at distal nodes; peduncle 0.1–0.6(–2) mm. |
Euphorbia radians |
Euphorbia humistrata |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–late summer. |
Habitat | Pinyon-juniper woodlands, oak savannas, desert grasslands and scrub. | Stream and river banks, gravel bars, floodplains, pond edges, disturbed fields, railroads, roadsides. |
Elevation | 700–2500 m. (2300–8200 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
AZ; TX; Mexico
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AL; AR; FL; GA; IL; IN; KS; KY; LA; MO; MS; OH; OK; TN; TX; VA; WV
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Discussion | Euphorbia radians is widely distributed but scattered from the Sonoran and Chihuahuan deserts south to Oaxaca in Mexico. The species is distinct among species in sect. Poinsettia in the flora area in its precocious habit, often flowering before the leaves emerge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia humistrata is distributed throughout the Mississippi River valley and along other major river systems in the central and eastern United States. There are scattered reports of this species as a waif or as introduced farther north and/or east (for example, Minnesota, New York, North Carolina, Ontario, South Dakota, and Wisconsin), but the authors have not been able to verify these occurrences. Euphorbia humistrata is similar to E. maculata and is often confused with that species in herbaria. It can be distinguished from E. maculata by its tendency to root at the stem nodes, its longer styles, and its seeds that lack low transverse ridges and that are more bluntly angled. When growing side-by-side, E. humistrata has an overall less congested appearance and its cyathia are not as numerous or crowded as those of E. maculata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 324. | FNA vol. 12, p. 272. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Poinsettia radians | Chamaesyce humistrata |
Name authority | Bentham: Pl. Hartw., 8. (1839) | Engelmann: in A. Gray, Manual ed. 2, 386. (1856) |
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