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sun spurge

Hoover's sandmat, Hoover's spurge

Habit Herbs, perennial, with moniliform tuberous rootstock. Herbs, annual, with slender to slightly thickened taproot.
Stems

erect, 5–20(–30) cm, usually glabrous, occasionally puberulent;

branches ± straight.

prostrate, occasionally mat-forming, 4–10 cm, glabrous.

Leaves

alternate;

petiole 0–2 mm, glabrous or strigose;

blade linear-lanceolate to ovate or broadly elliptic, 25–50 × 3–20 mm, unlobed, base rounded (tapered to petiole), margins with few glandular teeth, strigillose, flat to revolute, apex acute, abaxial surface coarsely strigose, adaxial surface strigose-hirsute;

venation pinnate, midvein prominent.

opposite;

stipules distinct or connate at base, deeply laciniate into numerous subulate to filiform segments, 0.3–1.3 mm, glabrous;

petiole 0–0.5 mm, glabrous;

blade broadly ovate, broadly oblong or suborbiculate, 3–7 × 2–5.5 mm, base asymmetric, hemiamplexicaulous, margin sharply serrulate to denticulate, teeth whitish and occasionally setaelike at apex, apex obtuse, surfaces papillate, glabrous; 3-veined from base, usually only midvein conspicuous.

Involucre

broadly globose-cupulate, 1.7–2.1 × 2.2–2.5 mm, glabrous or puberulent;

involucral lobes divided into triangular segments;

glands 1–4(–5), white, sessile and broadly attached, 1.1 × 1.4 mm, opening oblong, glabrous;

appendages absent.

campanulate, 1.2–1.5 × 1.5–2.2 mm, papillate;

glands 4, yellow to reddish, subcircular to oblong, 0.4–0.5 × 0.5–0.7 mm;

appendages white to pink, 0.3–0.9 × 0.7–1.6 mm, divided into 4–6 narrowly triangular, acute segments, these occasionally 2-fid, margins entire.

Staminate flowers

20–25.

25–35.

Pistillate flowers

ovary glabrous or puberulent, styles 3–4 mm, 2-fid 1/2 to nearly entire length.

ovary glabrous;

styles 1.8–2.6 mm, unbranched, filiform.

Capsules

depressed-globose, 3.8–5 × 4–5 mm, 3-lobed, glabrous or puberulent;

columella 3.6–4.5 mm.

oblate, 1.5–1.7 × 1.8–2.2 mm, glabrous;

columella 1.2–1.4 mm.

Seeds

white, mottled brown to gray, ellipsoid, rounded in cross section, 4–4.6 × 2.4–3.2 mm, smoothly and broadly pitted or grooved;

caruncle 0.1 mm.

gray to light brown, ovoid, inconspicuously 4-angled in cross section, 1.2–1.6 × 1–1.1 mm, rugose.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched (often highly condensed);

pleiochasial bracts 6–8(–10), as tight involucrate whorl, wholly white to pale pink or red, usually narrower than distal leaves;

dichasial bracts linear and highly reduced.

Cyathia

peduncle 2–5.5 mm.

solitary at distal nodes;

peduncle 0.5 mm, stout.

Euphorbia radians

Euphorbia hooveri

Phenology Flowering and fruiting spring–summer. Flowering and fruiting summer–fall.
Habitat Pinyon-juniper woodlands, oak savannas, desert grasslands and scrub. Drying mudflats of vernal pools in grasslands and woodlands.
Elevation 700–2500 m. (2300–8200 ft.) 20–200 m. (100–700 ft.)
Distribution
from FNA
AZ; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Euphorbia radians is widely distributed but scattered from the Sonoran and Chihuahuan deserts south to Oaxaca in Mexico. The species is distinct among species in sect. Poinsettia in the flora area in its precocious habit, often flowering before the leaves emerge.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia hooveri is federally listed as threatened; its populations are being affected severely by habitat loss and the invasion of exotic species. The species is endemic to vernal pools in six counties in the Central Valley of California. Molecular data show that E. hooveri is a hybrid species, closely related to E. albomarginata and E. serpens (Y. Yang and P. E. Berry 2011).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 324. FNA vol. 12, p. 271.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Poinsettia radians Chamaesyce hooveri
Name authority Bentham: Pl. Hartw., 8. (1839) L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 9. (1940)
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