Euphorbia radians |
Euphorbia brachycera |
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sun spurge |
horned spurge, shorthorn spurge |
|
Habit | Herbs, perennial, with moniliform tuberous rootstock. | Herbs, perennial, with thick rootstock. |
Stems | erect, 5–20(–30) cm, usually glabrous, occasionally puberulent; branches ± straight. |
erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
Leaves | alternate; petiole 0–2 mm, glabrous or strigose; blade linear-lanceolate to ovate or broadly elliptic, 25–50 × 3–20 mm, unlobed, base rounded (tapered to petiole), margins with few glandular teeth, strigillose, flat to revolute, apex acute, abaxial surface coarsely strigose, adaxial surface strigose-hirsute; venation pinnate, midvein prominent. |
petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
Involucre | broadly globose-cupulate, 1.7–2.1 × 2.2–2.5 mm, glabrous or puberulent; involucral lobes divided into triangular segments; glands 1–4(–5), white, sessile and broadly attached, 1.1 × 1.4 mm, opening oblong, glabrous; appendages absent. |
turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
Staminate flowers | 20–25. |
10–15. |
Pistillate flower(s) | ovary glabrous or puberulent, styles 3–4 mm, 2-fid 1/2 to nearly entire length. |
ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
Capsules | depressed-globose, 3.8–5 × 4–5 mm, 3-lobed, glabrous or puberulent; columella 3.6–4.5 mm. |
depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
Seeds | white, mottled brown to gray, ellipsoid, rounded in cross section, 4–4.6 × 2.4–3.2 mm, smoothly and broadly pitted or grooved; caruncle 0.1 mm. |
light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched (often highly condensed); pleiochasial bracts 6–8(–10), as tight involucrate whorl, wholly white to pale pink or red, usually narrower than distal leaves; dichasial bracts linear and highly reduced. |
arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
Cyathia | peduncle 2–5.5 mm. |
peduncle 0.4–1 mm. |
2n | = 28. |
|
Euphorbia radians |
Euphorbia brachycera |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–summer. |
Habitat | Pinyon-juniper woodlands, oak savannas, desert grasslands and scrub. | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. |
Elevation | 700–2500 m. (2300–8200 ft.) | 1200–3200 m. (3900–10500 ft.) |
Distribution |
AZ; TX; Mexico
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AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
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Discussion | Euphorbia radians is widely distributed but scattered from the Sonoran and Chihuahuan deserts south to Oaxaca in Mexico. The species is distinct among species in sect. Poinsettia in the flora area in its precocious habit, often flowering before the leaves emerge. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 324. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Poinsettia radians | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus |
Name authority | Bentham: Pl. Hartw., 8. (1839) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) |
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