Euphorbia prostrata |
Euphorbia helleri |
|
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blue weed, ground spurge, prostrate sandmat, prostrate spurge or sandmat |
Heller's spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | prostrate to decumbent, usually not mat-forming, 10–30 cm, crisped-villous to glabrate (proximally). |
ascending, branched proximally, 15–30 cm, glabrous. |
Leaves | opposite; stipules connate (lower side) or distinct (upper side), triangular-subulate, often lacerate distally, 0.5–1 mm, short crisped-villous to glabrate; petiole 0.5–1.5 mm, usually short crisped-villous, sometimes wooly at distal nodes; blade broadly elliptic to elliptic-oblong, ovate-spatulate, or ovate, 3–11(–15) × 3–6(–8) mm, base slightly asymmetric, rounded to slightly cordate and oblique, margins serrulate at least in distal 1/2, sometimes obscurely so, apex obtuse, abaxial surface finely crisped-villous, adaxial surface usually glabrous or glabrate, sometimes sparsely crisped-villous; 3-veined from base. |
petiole usually 1–3 mm, absent distally; blade spatulate, 6–15 × 3–5 mm, base broadly attenuate, margins entire, apex usually obtuse to rounded, sometimes retuse, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | obconic, 0.6–0.9 × 0.5 mm, crisped-villous or glabrous; glands 4, reddish, oval to oblong, 0.1 × 0.1–0.2 mm; appendages white to pink, rudimentary, 0–0.2 mm, distal margin entire or irregularly scalloped. |
campanulate, 0.9–1.2 × 0.7–1 mm, glabrous; glands 4, elliptic, 0.1–0.2 × 0.3–0.4 mm; horns slightly divergent, 0.1–0.2 mm. |
Staminate flowers | 3–6. |
8–10. |
Pistillate flowers | ovary densely crisped-villous; styles 0.1 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.4–0.5 mm, 2-fid. |
Capsules | broadly ovoid, 1.2–2 × 1.4–1.5 mm, crisped-villous along keels and toward base, often glabrous between keels; columella 1–1.2 mm. |
depressed-globose, 2.1–2.5 × 2.5–3 mm, 3-lobed; cocci flattened, smooth, glabrous; columella 1.2–1.7 mm. |
Seeds | white but with barely concealed brown surface beneath, ovoid, sharply 4-angled in cross section, abaxial faces plane to convex, adaxial faces concave, 0.8–1.1 × 0.5–0.7 mm, with several narrow, sharp, slightly irregular, transverse ridges. |
white to light gray, ovoid, 1.4–1.6 × 1–1.2 mm, smooth; caruncle 2-lobed, thin, 0.4 × 0.7 mm. |
Cyathia | solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 1–2 mm. |
peduncle 0.2–0.4 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 1–5 times 2-branched; pleiochasial bracts oblong, similar in size to distal leaves; dichasial bracts distinct, orbiculate-ovate to nearly reniform, subpandurate, base broadly cuneate to truncate, margins entire, apex obtuse, mucronate; axillary cymose branches 0–5. |
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2n | = 18. |
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Euphorbia prostrata |
Euphorbia helleri |
|
Phenology | Flowering and fruiting early spring–fall. | Flowering and fruiting late winter–spring. |
Habitat | Disturbed areas, fields, gardens, sidewalks, sandy places, ballast piles. | Forests, stream banks, roadsides, shaded areas with sandy, calcareous soils. |
Elevation | 0–1400 m. (0–4600 ft.) | 0–50 m. (0–200 ft.) |
Distribution |
AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NM; OH; OK; PA; SC; SD; TN; TX; UT; VA; WV; WY; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Pacific Islands, Australia]
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LA; TX; Mexico (Nuevo León) |
Discussion | Euphorbia prostrata is native to tropical America and possibly into the southern part of the flora area. It is likely adventive throughout most of the northern part of its range. It is widely naturalized throughout much of the rest of the tropics. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The smooth, white to light gray seeds of Euphorbia helleri easily distinguish it from other annual members of subg. Esula in North America. Collections of E. helleri have been made near Brownsville, Texas, and thus it is possible that the species occurs in northern Tamaulipas, Mexico. The Louisiana record (Webster Parish) likely represents introduced plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 285. | FNA vol. 12, p. 303. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce prostrata | Tithymalus helleri |
Name authority | Aiton: Hort. Kew. 2: 139. (1789) | Millspaugh: Bot. Gaz. 26: 268, fig. [p. 270]. (1898) |
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