Euphorbia polygonifolia |
Euphorbia trichotoma |
|
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dune spurge, euphorbe à feuilles de renouée, seaside sandmat, seaside spurge |
sand-dune spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, perennial, with thick rootstock. |
Stems | usually prostrate, occasionally ascending, 5–30 cm, glabrous. |
erect, branched, 15–40 cm, glabrous. |
Leaves | opposite; stipules usually distinct, occasionally connate basally (distal portion of stem), triangular-subulate, entire or divided, 0.8–1.5 mm, glabrous; petiole 1–3 mm, glabrous; blade oblong, linear-oblong, or linear-lanceolate, 5–16 × 2–4 mm, base slightly asymmetric, obtuse or subcordate, margins entire, apex obtuse, often mucronulate, surfaces uniformly green or reddish tinged, glabrous; venation obscure. |
petiole absent; blade obovate to oblanceolate, 5–20 × 2–7 mm, base broadly attenuate, margins entire, apex acute to obtuse, surfaces glabrous; venation pinnate, very obscure, midvein prominent. |
Involucre | obconic-campanulate, 1.2–1.7 × 1–1.4 mm, glabrous; glands 4, occasionally rudimentary, green-yellow to tan or orange-tinged, occasionally shortly stipitate, usually broadly oval to subcircular, sometimes figure eight-shaped, shallowly cupped, 0.1–0.3 × 0.2–0.4 mm; appendages absent or rudimentary. |
hemispheric, 1–1.5 × 1.4–2 mm, glabrous; glands 5, elliptic to slightly crescent-shaped, 0.4–0.6 × 0.9–1.1 mm; horns divergent, 0.1–0.2 mm. |
Staminate flowers | 5–14. |
8–10. |
Pistillate flowers | ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.2–0.4 mm, 2-fid. |
Capsules | broadly ovoid, 3–3.5(–4) × (2–)2.4–3 mm, glabrous; columella 2–3 mm. |
depressed-ovoid, 1.1–1.5 × 2–3 mm, strongly 3-lobed; cocci rounded, minutely papillate, sometimes appearing smooth, glabrous; columella 1.6–1.9 mm. |
Seeds | ashy white, wedge-shaped to slightly ovoid, weakly dorsiventrally compressed and elliptic-terete to bluntly subangled in cross section, back strongly rounded, face slightly rounded, (2–)2.2–2.8 × 1.6–1.9 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
white, ovoid-globose, 1.8–2 × 1.6–1.8 mm, smooth; caruncle ± rounded and flattened, 0.3 × 0.3 mm. |
Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 0.5–5 mm. |
peduncle 0.7–1 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each usually 1–2 times 3-branched, sometimes unbranched; pleiochasial bracts similar in shape and size to distal leaves, alternate; dichasial bracts distinct, obovate to oblanceolate, base broadly attenuate, margins entire, apex acute to obtuse; axillary cymose branches 0. |
|
Euphorbia polygonifolia |
Euphorbia trichotoma |
|
Phenology | Flowering and fruiting early summer–fall. | Flowering and fruiting year-round (primarily spring–summer). |
Habitat | Sandy maritime and freshwater beaches and foredunes. | Coastal beaches, sand dunes, thickets. |
Elevation | 0–200 m. (0–700 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
CT; DE; FL; GA; IL; IN; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; SC; VA; WI; NB; NS; ON; PE; QC
|
FL; Mexico; Central America; West Indies |
Discussion | Euphorbia polygonifolia is native to coastal beaches and dunes along the Atlantic Ocean from the maritime provinces of Canada south to northern Florida. The species also occurs disjunctly along the shores of the North American Great Lakes. Euphorbia polygonifolia was native to Quebec but is now considered extirpated from the province. It has apparently been introduced in Europe, but it is unclear if it has persisted there (L. C. Wheeler 1941). Immature individuals of this species can be somewhat difficult to distinguish from E. bombensis where their ranges overlap (Virginia to northern Florida). Where they occur together, E. polygonifolia tends to be a pioneer species on the upper beach and foredune front, whereas E. bombensis tends to inhabit areas behind the foredune (R. D. Porcher and D. A. Rayner 2002). Euphorbia polygonifolia can be distinguished also by its larger capsules and larger, wedge-shaped to slightly ovoid seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia trichotoma is found in coastal peninsular Florida from Hillsborough County south to Key West. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 284. | FNA vol. 12, p. 311. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce polygonifolia | Galarhoeus trichotomus, Tithymalus trichotomus |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 48; 2(qto.): 60. (1817) |
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