Euphorbia polygonifolia |
Euphorbia jaegeri |
|
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dune spurge, euphorbe à feuilles de renouée, seaside sandmat, seaside spurge |
orocopia mountains spurge |
|
Habit | Herbs, annual, with taproot. | Shrubs, with woody rootstock. |
Stems | usually prostrate, occasionally ascending, 5–30 cm, glabrous. |
ascending, diffusely and intricately branched, 15–25 cm, usually puberulent to shortly hirsute, sometimes glabrate, bark grayish. |
Leaves | opposite; stipules usually distinct, occasionally connate basally (distal portion of stem), triangular-subulate, entire or divided, 0.8–1.5 mm, glabrous; petiole 1–3 mm, glabrous; blade oblong, linear-oblong, or linear-lanceolate, 5–16 × 2–4 mm, base slightly asymmetric, obtuse or subcordate, margins entire, apex obtuse, often mucronulate, surfaces uniformly green or reddish tinged, glabrous; venation obscure. |
opposite; stipules distinct or connate, subulate, 0.3–0.5 mm, puberulent; petiole 0.7–1.1 mm, puberulent to shortly hirsute; blade ovate or elliptic, 3–9 × 1.5–5 mm, base symmetric to slightly asymmetric, rounded to cuneate, margins entire, apex usually obtuse, sometimes acute, surfaces puberulent to shortly hirsute; 3-veined from base, often only midvein conspicuous. |
Involucre | obconic-campanulate, 1.2–1.7 × 1–1.4 mm, glabrous; glands 4, occasionally rudimentary, green-yellow to tan or orange-tinged, occasionally shortly stipitate, usually broadly oval to subcircular, sometimes figure eight-shaped, shallowly cupped, 0.1–0.3 × 0.2–0.4 mm; appendages absent or rudimentary. |
obconic to campanulate, 1.2–1.8 × 1.1–1.4 mm, puberulent to shortly hirsute; glands 4, yellow to pinkish, elliptic to oblong, 0.3 × 0.4–0.5 mm; appendages white to pink, 0.2–0.7 × 0.6–1.2 mm, irregularly divided from halfway to nearly base into 4–8 triangular to subulate segments, segments entire. |
Staminate flowers | 5–14. |
25–30. |
Pistillate flowers | ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
ovary canescent; styles 0.3–0.4 mm, 2-fid entire length. |
Capsules | broadly ovoid, 3–3.5(–4) × (2–)2.4–3 mm, glabrous; columella 2–3 mm. |
oblate, 1.7–2.3 × 1.8–2.7 mm, puberulent; columella 1.4–2 mm. |
Seeds | ashy white, wedge-shaped to slightly ovoid, weakly dorsiventrally compressed and elliptic-terete to bluntly subangled in cross section, back strongly rounded, face slightly rounded, (2–)2.2–2.8 × 1.6–1.9 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
tan to grayish, narrowly oblong-ovoid, ± 3–4-angled in cross section, 1.4–1.5 × 0.7–0.9 mm, irregularly dimpled or with faint transverse ridges that do not interrupt abaxial keel. |
Cyathia | solitary or in small, cymose clusters at distal nodes; peduncle 0.5–5 mm. |
solitary at distal nodes; peduncle 0.5–1.7 mm. |
Euphorbia polygonifolia |
Euphorbia jaegeri |
|
Phenology | Flowering and fruiting early summer–fall. | Flowering and fruiting fall–spring. |
Habitat | Sandy maritime and freshwater beaches and foredunes. | Desert scrub, hillsides, arroyos, primarily in rock crevices. |
Elevation | 0–200 m. (0–700 ft.) | 600–900 m. (2000–3000 ft.) |
Distribution |
CT; DE; FL; GA; IL; IN; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; SC; VA; WI; NB; NS; ON; PE; QC
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CA |
Discussion | Euphorbia polygonifolia is native to coastal beaches and dunes along the Atlantic Ocean from the maritime provinces of Canada south to northern Florida. The species also occurs disjunctly along the shores of the North American Great Lakes. Euphorbia polygonifolia was native to Quebec but is now considered extirpated from the province. It has apparently been introduced in Europe, but it is unclear if it has persisted there (L. C. Wheeler 1941). Immature individuals of this species can be somewhat difficult to distinguish from E. bombensis where their ranges overlap (Virginia to northern Florida). Where they occur together, E. polygonifolia tends to be a pioneer species on the upper beach and foredune front, whereas E. bombensis tends to inhabit areas behind the foredune (R. D. Porcher and D. A. Rayner 2002). Euphorbia polygonifolia can be distinguished also by its larger capsules and larger, wedge-shaped to slightly ovoid seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia jaegeri is known only from the Orocopia Mountains of Riverside County and the Bristol and Marble Mountains of San Bernardino County. The species is one of few shrubby species of sect. Anisophyllum in the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 284. | FNA vol. 12, p. 273. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce polygonifolia | |
Name authority | Linnaeus: Sp. Pl. 1: 455. (1753) | V. W. Steinmann & J. M. André: Aliso 30: 1, figs. 1–4. (2012) |
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