Euphorbia polycarpa |
Euphorbia trachysperma |
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smallseed sandmat |
San Pedro River sandmat |
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Habit | Herbs, annual, with slender taproot. | |
Stems | erect to ascending, 10–55 cm, glabrous. |
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Leaves | opposite; stipules distinct, narrowly triangular and often divided into 3–5 subulate to filiform segments, 0.8–1.9 mm, glabrous; petiole 1.2–3.1 mm, glabrous; blade oblong, elliptic, or ovate, 12–43 × 3–14 mm, base subsymmetric to asymmetric, rounded, attenuate, or with one side hemicordate and other side rounded, margins usually serrulate, rarely entire on some leaves or portion of blade, apex obtuse to acute, surfaces often with red blotch in center, glabrous; pinnately veined, often only midvein conspicuous. |
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Involucre | oblong, campanulate, or funnel-shaped, 1.5–2.6 × 1.1–2.3 mm, glabrous; glands 4, green to yellowish or tinged with red, subcircular to oblong, 0.4–0.7 × 0.5–1 mm; appendages absent or white, often forming narrow rim or oblong to ovate, 0–0.8 × 1.2 mm, distal margin entire. |
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Staminate flowers | 50–60. |
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Pistillate flowers | ovary glabrous; styles 0.5–0.6 mm, 2-fid 1/2 length. |
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Capsules | oblate, 2.4–3.2 × 3–3.5 mm, glabrous; columella 2.3–2.5 mm. |
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Seeds | light brown, ovoid, 4-angled in cross section, abaxial keel well developed, 1.9–2.3 × 1.3–1.4 mm, finely dimpled and papillate. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at distal nodes; peduncle 1.4–2.4 mm. |
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Euphorbia polycarpa |
Euphorbia trachysperma |
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Phenology | Flowering and fruiting late summer–early winter. | |
Habitat | Desert scrub, desert grasslands, mesquite woodlands. | |
Elevation | 200–1200 m. (700–3900 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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AZ; Mexico (Baja California Sur, Sonora) |
Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia trachysperma may be expected in extreme southwestern New Mexico given the close proximity of some collections in southeastern Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 291. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | Chamaesyce trachysperma |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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