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smallseed sandmat
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mat spurge, narrow-seed spurge, slimseed sandmat
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Herbs, annual, with taproot. |
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prostrate, often mat-forming, occasionally with ascending tips, 5–45 cm, densely and evenly pilose to lanate. |
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opposite; stipules distinct or connate basally on one side of stem, entire or irregularly toothed or fringed, 0.5–1.2 mm, pilose to lanate; petiole 0.3–1.5 mm, pilose to lanate; blade usually oblong to oblong-obovate, occasionally nearly circular, 3–10(–15) × 2–5(–10) mm, base asymmetric, one side usually angled or rounded and other truncate-auriculate, margins minutely or conspicuously serrulate at least toward apex, apex usually broadly rounded to broadly acute, occasionally emarginate, abaxial surface often ± lighter green and without reddish spot, both surfaces sparsely to moderately pilose to lanate; 3-veined from base or venation obscure. |
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obconic, 0.7–1 × 0.4–0.6 mm, moderately to densely pilose to lanate; glands 4, reddish, ± unequal, oblong, 0.1 × 0.1–0.3 mm; appendages white to strongly pinkish or reddish tinged, often unequal, sometimes 1 to all absent, 0–0.3 × 0–0.4 mm, 3-lobed or rudimentarily 1-lobed, distal margin crenate. |
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3–9. |
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ovary pilose to villous, hairs occasionally slightly appressed; styles 0.2–0.5 mm, unbranched or inconspicuously notched at tip. |
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ovoid, 1.6–2.3 × 1.4–1.5 mm, moderately to densely villous with hairs usually slightly appressed, pubescence often concentrated on proximal 1/2 or along lobes; columella 1.5–2 mm. |
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light to dark brown, usually mottled, sometimes with thin, white coating, often wearing away irregularly, narrowly oblong-ovoid to ellipsoid, 3–4-angled in cross section, 1–1.5 × 0.5–0.6 mm, with short, irregularly interrupted furrows, appearing partially and irregularly few-ridged. |
polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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(1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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reported two of these for the United States: var. hirtella and var. polycarpa. |
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hirtella, as the name suggests, was applied to hairy plants. |
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, it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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solitary at leaf nodes or in small, cymose clusters on congested, axillary branches; peduncle 0.7–2.5 mm. |
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Flowering and fruiting midsummer–early fall. |
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Open disturbed areas, rocky slopes. |
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100–2100 m. (300–6900 ft.) |
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AZ; CA; NV; nw Mexico
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AZ; CO; IA; KS; MO; ND; NE; NM; OK; SD; TX; WY; Mexico (Chihuahua, Durango, San Luis Potosí)
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Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia stictospora has been recorded from New York, but this disjunct occurrence likely represents a waif or misidentification. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 283. |
FNA vol. 12, p. 289. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
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| Chamaesyce polycarpa |
Chamaesyce stictospora |
| Bentham: Bot. Voy. Sulphur, 50. (1844) |
Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) |
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