Euphorbia polycarpa |
Euphorbia setiloba |
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smallseed sandmat |
fringe spurge, shaggy spurge, Yuma sandmat |
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Habit | Herbs, annual, with slender taproot. | |
Stems | prostrate, mat-forming, 5–50 cm, villous with glistening glandular hairs. |
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Leaves | opposite; stipules distinct, filiform, rudimentary to 0.2 mm, glabrous or sparsely villous with glistening glandular hairs; petiole 0.5–1.5 mm, villous; blade oblong, ovate, or elliptic, 3–7 × 2–4 mm, base asymmetric, rounded, margins entire, apex obtuse, surfaces villous; weakly 3-veined from base, commonly only midvein conspicuous. |
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Involucre | campanulate or urceolate, 0.7–1 × 0.5–0.8 mm, villous; glands 4, red to pink, oblong to slightly reniform, 0.1–0.2 × 0.2–0.3 mm; appendages white to pink, deeply incised into 3–6 triangular to subulate, attenuate, acute segments, 0.3–0.6 × 0.6–1 mm, segments entire. |
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Staminate flowers | 3–7. |
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Pistillate flowers | ovary villous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
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Capsules | subglobose to ovoid, 1–1.2 mm diam., villous; columella 0.9–1.1 mm. |
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Seeds | pink to light gray, narrowly ovoid, 4-angled in cross section, 0.8–1 × 0.5–0.6 mm, dimpled or with faint transverse ridges that do not pass through abaxial keel. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at distal nodes, nodes often congested toward tips of branches; peduncle 0.2–1.6 mm. |
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Euphorbia polycarpa |
Euphorbia setiloba |
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Phenology | Flowering nearly year-round in response to sufficient moisture. | |
Habitat | Desert scrub, blackbrush scrub, Joshua tree woodlands, grasslands, often in sandy areas. | |
Elevation | 20–1600 m. (100–5200 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Durango, Sinaloa, Sonora)
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 288. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | Chamaesyce setiloba |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Engelmann: in War Department [U.S.], Pacif. Railr. Rep. 5(2): 364. (1857) |
Web links |