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smallseed sandmat
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revolute spurge, threadstem sandmat, threadstem spurge
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Herbs, annual, with slender taproot. |
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erect, 5–25 cm, glabrous. |
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opposite; stipules distinct, subulate-filiform, entire, 0.5–0.9 mm, glabrous; petiole 0.6–1.1 mm, glabrous; blade linear, 6–27 × 0.6–1.2 mm, base symmetric, attenuate, margins entire, revolute, apex acute, sometimes mucronate, surfaces glabrous; only midvein conspicuous. |
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obconic to campanulate, 0.7–0.9 × 0.5–0.7 mm, glabrous; glands 4, pink to dark purple, nearly circular to oblong or reniform, 0.1 × 0.1–0.2 mm; appendages white, oblong, nearly circular, ovate, deltate, or forming thin margin around gland, rarely absent, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin entire. |
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5–10. |
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ovary glabrous; styles 0.2–0.3 mm, unbranched. |
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ovoid to subglobose, 1.5–1.8 × 1.6–1.8 mm, glabrous; columella 1.1–1.5 mm. |
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whitish, brick red, light gray, or light brown, narrowly to broadly ovoid, 4-angled in cross section, 0.9–1.4 × 0.7–1 mm, nearly smooth, rugulose, with faint transverse ridges, or with 2–3 well-defined transverse ridges separated by shallow depressions. |
polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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(1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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reported two of these for the United States: var. hirtella and var. polycarpa. |
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hirtella, as the name suggests, was applied to hairy plants. |
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, it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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solitary at distal nodes; peduncles absent or to 1.5 mm. |
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Flowering and fruiting summer–late fall. |
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Desert scrub, sagebrush scrub, juniper woodlands, Joshua tree-pinyon pine woodlands, oak woodlands, grasslands, chaparral, pine-oak forests. |
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600–2500 m. (2000–8200 ft.) |
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AZ; CA; NV; nw Mexico
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AZ; CA; CO; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Sonora, Zacatecas)
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Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia revoluta is a distinctive, easily recognizable species by virtue of the combination of unbranched styles and linear leaves with symmetric bases. The species ranges through northern Mexico into the southwestern United States and is composed of three well-marked, geographically distinct races that can be distinguished by their seeds. The first of these races occurs primarily in pine-oak forest of the Sierra Madre Occidental of Mexico (Chihuahua and Sonora) and barely enters the flora area in the Huachuca Mountains of southeastern Arizona; the seeds are brick red and nearly smooth. The second race is widespread in the Chihuahuan Desert from northern Mexico to southeastern New Mexico and southwestern Texas; its seeds are whitish to light gray and possess two or three well-defined transverse ridges separated by shallow depressions. The third race corresponds to the type collection and is characterized by grayish white to light brown seeds that are rugulose or with faint transverse ridges; it occurs in northwestern Mexico (Baja California and Sonora) and throughout the arid southwestern United States. It is probable that further study will justify the taxonomic segregation of these races as either distinct species or subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 283. |
FNA vol. 12, p. 286. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
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| Chamaesyce polycarpa |
Chamaesyce revoluta |
| Bentham: Bot. Voy. Sulphur, 50. (1844) |
Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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