Euphorbia polycarpa |
Euphorbia marginata |
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smallseed sandmat |
euphorbe marginée, mountain snow spurge, smoke-on-the-prairie, snow-on-the-mountain, variegated spurge, whitemargined spurge |
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Habit | Herbs, annual, with taproot. | |
Stems | erect, unbranched or branched, 30–85(–150) cm, pilose or glabrous. |
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Leaves | alternate; stipules 0.1–0.3 mm; petiole 0.2–3 mm, glabrous or minutely pilose; blade broadly ovate to elliptic, 32–62(–82) × 18–28(–52) mm, base rounded to cuneate, margins entire, often white on distal leaves, apex acute, rarely mucronate, surfaces glabrous; venation obscure, only midvein conspicuous. |
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Involucre | campanulate, 2.2–3.5 × 1.3–2.3 mm, margin between glands deeply divided into fringe of fimbriate lobes, pilose; glands 4–5, green to greenish yellow, reniform to subcircular, 0.7–1.1 × 1–1.6 mm; appendages white, orbiculate, 1.5–2.7 × 1.9–2.9(–3.6) mm, entire. |
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Staminate flowers | 30–70. |
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Pistillate flowers | ovary pilose; styles 1–2.5 mm, 2-fid 1/2–2/3 length. |
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Capsules | oblate, 3–5 × 3.5–7.5 mm, moderately to densely pilose; columella 3–4.1 mm. |
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Seeds | orange-tan to gray, ovoid, 3.7–3.9 × 3–3.3 mm, rugose, with 2 transverse ridges (one dark orange to brown, other inconspicuous); caruncle absent. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | in terminal pleiochasia, dichasial bracts narrowly elliptic to oblanceolate with conspicuous white margins; peduncle 1.8–2.7(–22) mm, densely pilose. |
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2n | = 56. |
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Euphorbia polycarpa |
Euphorbia marginata |
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Phenology | Flowering and fruiting summer–fall. | |
Habitat | Disturbed areas and grasslands. | |
Elevation | 0–1700 m. (0–5600 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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AR; CA; CO; CT; IA; IL; IN; KS; LA; MA; MD; MI; MN; MO; MT; NC; NE; NH; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; WI; WV; WY; MB; ON; QC; SK; c Mexico; s Mexico
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia marginata is native to the central United States. The type specimen was collected by Meriwether Lewis along the Yellowstone River in southern Montana in 1806, and it has been reported to be native as far south as Arizona, New Mexico, and Texas, and as far east as southern Minnesota, western Iowa, and Missouri (G. Yatskievych 1999–2013, vol. 2). It is presumably naturalized outside of this area. Euphorbia marginata is widely cultivated as an ornamental for its showy, white-margined distal leaves, and it can escape locally. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 248. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | E. bonplandii, Lepadena marginata |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Pursh: Fl. Amer. Sept. 2: 607. (1813) |
Web links |
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