FNA: Euphorbia polycarpa vs. Euphorbia lasiocarpa

	Euphorbia polycarpa	Euphorbia lasiocarpa
	smallseed sandmat	roadside sandmat
Habit		Herbs, annual or perennial, often robust, with slender to moderately thickened taproot.
Stems		erect to ascending, 30–100 cm, pilose to tomentose.
Leaves		opposite; stipules distinct when young, connate into deltate scale when older, erose to laciniate, with dark glands at margin or base, 0.5–1.3 mm, pilose; petiole 0.5–2 mm, glabrescent or pilose; blade ovate or oblong, sometimes slightly falcate, 8–46 × 3–21 mm, base asymmetric, obtuse to hemicordate, margins serrulate, apex obtuse or acute, surfaces often with red spot in center, pilose to sericeous; palmately 3–5(–7)-veined from base.
Involucre		obconic, 0.9–1.3 × 0.8–1.2 mm, pilose to sericeous; glands 4, yellow or pink, circular or oblong, 0.1–0.3 mm diam.; appendages white or pink, oblong, flabellate, or suborbiculate, 0.1–0.6 × 0.2–1 mm, entire or crenate.
Staminate flowers		15–25.
Pistillate flowers		ovary densely pilose to sericeous with yellowish hairs; styles 0.6–0.9 mm, 2-fid 1/2 length, filiform.
Capsules		subglobose to broadly ovoid, 1.7–2 mm diam., pilose to sericeous with yellowish hairs; columella 1.5–1.7 mm.
Seeds		dark reddish brown to almost black, plumply ovoid, 4-angled in cross section, 1.2–1.3 × 0.7–0.8 mm, with 2 inconspicuous rows of 3–5 shallow depressions separated by low ridges.
Euphorbia	polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California.
l	.
c	.
Wheeler	(1941) divided the species into seven varieties, the majority of which occur in Baja California Sur.
He	reported two of these for the United States: var. hirtella and var. polycarpa.
Variety	hirtella, as the name suggests, was applied to hairy plants.
However	, it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual.
Cyathia		in dense, usually terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia; peduncle 0.8–2.3 mm.

	Euphorbia polycarpa	Euphorbia lasiocarpa
Phenology		Flowering and fruiting year-round.
Habitat		Open disturbed areas, mostly along roadsides and railroad tracks.
Elevation		0–10 m. (0–0 ft.)
Distribution	AZ; CA; NV; nw Mexico [WildflowerSearch map] [BONAP county map]	FL; Mexico; Central America; South America; West Indies [Introduced in North America] [BONAP county map]
Discussion	Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Euphorbia lasiocarpa is similar to E. hypericifolia but is much more hairy on its stems and leaves. Euphorbia lasiocarpa is widespread throughout tropical America, but its precise native range in the New World is not clear. In the flora area, E. lasiocarpa is found in southern Florida, where it is likely introduced. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 12, p. 283.	FNA vol. 12, p. 275.
Parent taxa	Euphorbiaceae > Euphorbia > sect. Anisophyllum	Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana	E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa	E. polycarpa var. polycarpa
Synonyms	Chamaesyce polycarpa	Chamaesyce lasiocarpa
Name authority	Bentham: Bot. Voy. Sulphur, 50. (1844)	Klotzsch: Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(suppl. 1): 414. (1843)
Web links	Local floras: CA Local Web sites: CalFlora, CalPhotos WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Euphorbia lasiocarpa

smallseed sandmat

roadside sandmat

Habit

Herbs, annual or perennial, often robust, with slender to moderately thickened taproot.

Stems

erect to ascending, 30–100 cm, pilose to tomentose.

Leaves

opposite;

stipules distinct when young, connate into deltate scale when older, erose to laciniate, with dark glands at margin or base, 0.5–1.3 mm, pilose;

petiole 0.5–2 mm, glabrescent or pilose;

blade ovate or oblong, sometimes slightly falcate, 8–46 × 3–21 mm, base asymmetric, obtuse to hemicordate, margins serrulate, apex obtuse or acute, surfaces often with red spot in center, pilose to sericeous; palmately 3–5(–7)-veined from base.

Involucre

obconic, 0.9–1.3 × 0.8–1.2 mm, pilose to sericeous;

glands 4, yellow or pink, circular or oblong, 0.1–0.3 mm diam.;

appendages white or pink, oblong, flabellate, or suborbiculate, 0.1–0.6 × 0.2–1 mm, entire or crenate.

Staminate flowers

15–25.

Pistillate flowers

ovary densely pilose to sericeous with yellowish hairs;

styles 0.6–0.9 mm, 2-fid 1/2 length, filiform.

Capsules

subglobose to broadly ovoid, 1.7–2 mm diam., pilose to sericeous with yellowish hairs;

columella 1.5–1.7 mm.

Seeds

dark reddish brown to almost black, plumply ovoid, 4-angled in cross section, 1.2–1.3 × 0.7–0.8 mm, with 2 inconspicuous rows of 3–5 shallow depressions separated by low ridges.

Euphorbia

polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California.

Wheeler

(1941) divided the species into seven varieties, the majority of which occur in Baja California Sur.

reported two of these for the United States: var. hirtella and var. polycarpa.

Variety

hirtella, as the name suggests, was applied to hairy plants.

However

, it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual.

Cyathia

in dense, usually terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia;

peduncle 0.8–2.3 mm.

Euphorbia polycarpa

Euphorbia lasiocarpa

Phenology

Flowering and fruiting year-round.

Habitat

Open disturbed areas, mostly along roadsides and railroad tracks.

Elevation

0–10 m. (0–0 ft.)

Distribution

AZ; CA; NV; nw Mexico

[WildflowerSearch map]

[BONAP county map]

FL; Mexico; Central America; South America; West Indies [Introduced in North America]

[BONAP county map]

Discussion

Varieties 5 (1 in the flora).

Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia lasiocarpa is similar to E. hypericifolia but is much more hairy on its stems and leaves. Euphorbia lasiocarpa is widespread throughout tropical America, but its precise native range in the New World is not clear. In the flora area, E. lasiocarpa is found in southern Florida, where it is likely introduced.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 12, p. 283.

FNA vol. 12, p. 275.

Parent taxa

Euphorbiaceae > Euphorbia > sect. Anisophyllum

Sibling taxa

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana

Subordinate taxa

E. polycarpa var. polycarpa

Synonyms

Chamaesyce polycarpa

Chamaesyce lasiocarpa

Name authority

Bentham: Bot. Voy. Sulphur, 50. (1844)

Klotzsch: Nov. Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19(suppl. 1): 414. (1843)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America

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Euphorbia polycarpa

Euphorbia lasiocarpa

Euphorbia polycarpa

Euphorbia lasiocarpa