Euphorbia polycarpa |
Euphorbia ipecacuanhae |
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smallseed sandmat |
American ipecac, Carolina ipecac, ipecac spurge |
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Habit | Herbs, perennial, with deep, stout rootstock. | |
Stems | decumbent or slightly ascending, branched, often densely clumped, previous year's dead stems not persistent, 17–27 cm, usually glabrous, rarely sparsely villous. |
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Leaves | alternate; stipules 0.1–0.2 mm; petiole (0–)1–2 mm, glabrous; blade usually linear, obovate, or oblanceolate to orbiculate, rarely filiform, proximal greatly reduced, scalelike, 15–70 × 1.5–13 mm gradually smaller proximally, base cuneate, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous; venation occasionally obscure on smaller leaves, midvein conspicuous. |
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Involucre | yellow or yellow-green, hemispheric, 1–1.2 × 2–2.4 mm, glabrous; glands 5, yellow or yellow-green, obovate or elliptic, 0.7–0.8 × 1–1.2 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
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Staminate flowers | 10–20. |
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Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
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Capsules | globose, 2.3–3.4 × 3.5–4.2 mm, glabrous; columella 3–3.1 mm. |
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Seeds | white or brown, ovoid, 2.3–2.5 × 1.5 mm, angular, with 5 longitudinal ridges, shallowly and irregularly pitted; caruncle absent. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 10–50(–70) mm, glabrous. |
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Euphorbia polycarpa |
Euphorbia ipecacuanhae |
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Phenology | Flowering and fruiting early spring–early summer. | |
Habitat | Pine and pine-oak savannas, pine-oak sand hills, turkey oak scrub, open sand habitats. | |
Elevation | 0–150 m. (0–500 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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CT; DC; DE; GA; MD; NC; NJ; NY; PA; SC; VA
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The vegetative stems of Euphorbia ipecacuanhae are often quite short in proportion to the dichasial or pleiochasial branches, thus superficially plants often appear to have mostly opposite leaves. However, careful examination of the base of the plant will reveal alternate leaves. The leaves are extremely variable in both shape and coloration, and the variation can be pronounced within a population or even on a single plant. M. J. Huft (1979) did not recognize infraspecific taxa within E. ipecacuanhae, and his treatment is followed here. This species is found on the Atlantic coastal plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 247. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | E. arundelana, Tithymalopsis ipecacuanhae |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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