Euphorbia polycarpa |
Euphorbia inundata |
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smallseed sandmat |
Florida pineland spurge |
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Habit | Herbs, perennial, with thickened rootstock. | |||||
Stems | erect or ascending, 15–40 cm. |
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Leaves | petiole absent or indistinct, blade linear to linear-elliptic or lanceolate, (25–)30–60(–115) × 1.5–14(–15) mm, chartaceous, base attenuate, apex acuminate, acute, or rounded and mucronate; only midvein evident. |
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Involucre | obconic or campanulate, 1.3–3.2 × 1.6–3.6 mm, lobes ovate, oblong, or nearly rectangular, 0.6–1 mm, laciniate-ciliate; glands red to greenish, oblong or nearly circular, 0.8–1 × 1.1–2 mm, distal margins crenulate-erose, undulate, or entire. |
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Staminate flowers | 20–25. |
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Pistillate flowers | gynophore exserted 2.6–6.1 mm, calyxlike lobes triangular to subulate, 0.6–1.4 mm; styles connate 1/8 length, 1.3–2.2 mm. |
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Capsules | oblate-ovoid, 5.1–6.1 × 6.9–8.6 mm, 3-lobed; columella 3.9–4.7 mm. |
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Seeds | brown to blackish, ovoid-globose, circular or faintly 3- or 4-angled in cross section, 2.9–3.4 × 2.7–3 mm, smooth, base flattened or rounded, apex rounded or with inconspicuous blunt point. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathial | arrangement: terminal pleiochasial branches 2–3, 5–18 cm, 3–8 times 2-branched; pleiochasial bracts linear-lanceolate or narrowly ovate, 25–49 × 5–9 mm, margins entire, apex acute or acuminate; dichasial bracts ovate or lanceolate, 6–21 × 2–6 mm, margins entire, apex acute or acuminate; axillary cymose branches 1–5. |
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Cyathia | peduncle 6–24 mm (often exceeding dichasial bracts). |
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Euphorbia polycarpa |
Euphorbia inundata |
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Distribution |
AZ; CA; NV; nw Mexico
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AL; FL; GA; MS |
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia inundata has a disjunct distribution in the southeastern United States. Allopatric, narrow-leaved populations from the west-central Florida peninsula are segregated as var. garrettii (E. L. Bridges and S. L. Orzell 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 314. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Nummulariopsis | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce polycarpa | Galarhoeus inundatus | ||||
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Torrey ex Chapman: Fl. South. U.S., 402. (1860) | ||||
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