Euphorbia polycarpa |
Euphorbia hooveri |
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smallseed sandmat |
Hoover's sandmat, Hoover's spurge |
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Habit | Herbs, annual, with slender to slightly thickened taproot. | |
Stems | prostrate, occasionally mat-forming, 4–10 cm, glabrous. |
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Leaves | opposite; stipules distinct or connate at base, deeply laciniate into numerous subulate to filiform segments, 0.3–1.3 mm, glabrous; petiole 0–0.5 mm, glabrous; blade broadly ovate, broadly oblong or suborbiculate, 3–7 × 2–5.5 mm, base asymmetric, hemiamplexicaulous, margin sharply serrulate to denticulate, teeth whitish and occasionally setaelike at apex, apex obtuse, surfaces papillate, glabrous; 3-veined from base, usually only midvein conspicuous. |
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Involucre | campanulate, 1.2–1.5 × 1.5–2.2 mm, papillate; glands 4, yellow to reddish, subcircular to oblong, 0.4–0.5 × 0.5–0.7 mm; appendages white to pink, 0.3–0.9 × 0.7–1.6 mm, divided into 4–6 narrowly triangular, acute segments, these occasionally 2-fid, margins entire. |
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Staminate flowers | 25–35. |
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Pistillate flowers | ovary glabrous; styles 1.8–2.6 mm, unbranched, filiform. |
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Capsules | oblate, 1.5–1.7 × 1.8–2.2 mm, glabrous; columella 1.2–1.4 mm. |
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Seeds | gray to light brown, ovoid, inconspicuously 4-angled in cross section, 1.2–1.6 × 1–1.1 mm, rugose. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at distal nodes; peduncle 0.5 mm, stout. |
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Euphorbia polycarpa |
Euphorbia hooveri |
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Phenology | Flowering and fruiting summer–fall. | |
Habitat | Drying mudflats of vernal pools in grasslands and woodlands. | |
Elevation | 20–200 m. (100–700 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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CA |
Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia hooveri is federally listed as threatened; its populations are being affected severely by habitat loss and the invasion of exotic species. The species is endemic to vernal pools in six counties in the Central Valley of California. Molecular data show that E. hooveri is a hybrid species, closely related to E. albomarginata and E. serpens (Y. Yang and P. E. Berry 2011). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 271. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | Chamaesyce hooveri |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 9. (1940) |
Web links |