Euphorbia polycarpa |
Euphorbia garberi |
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smallseed sandmat |
Garber's sandmat |
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Habit | Herbs, perennial, with moderately thickened rootstock. | |
Stems | ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent. |
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Leaves | opposite; stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose; petiole 0.5–1.1 mm, pilose; blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous. |
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Involucre | turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose; glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm; appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate. |
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Staminate flowers | 8–20. |
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Pistillate flowers | ovary pilose; styles 0.6–0.7 mm, 2-fid 1/2 length. |
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Capsules | subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose; columella 1.1–1.6 mm. |
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Seeds | gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at nodes of short, axillary branches; peduncle 0.2–0.4 mm. |
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Euphorbia polycarpa |
Euphorbia garberi |
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Phenology | Flowering and fruiting year-round. | |
Habitat | Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands. | |
Elevation | 0–10 m. (0–0 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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FL |
Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 268. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883) |
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