Euphorbia polycarpa |
Euphorbia bilobata |
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smallseed sandmat |
black-seed spurge |
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Habit | Herbs, annual, with slender taproot. | |
Stems | erect, branched, 10–35 cm, glabrous or strigillose (especially when young and around nodes). |
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Leaves | opposite proximally, alternate distally; stipules 0.1–0.2 mm; petiole 1–4(–6) mm, glabrous, sericeous or strigillose; blade linear to narrowly elliptic, 8–52 × 2–7 mm, base attenuate, margins entire, ciliate-strigose, apex acute, abaxial surface sparsely strigillose to sericeous, adaxial surface usually glabrous; venation obscure, only midvein conspicuous. |
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Involucre | obconic, 0.9–1.5 × 0.9–1.3 mm, strigillose to pilose; glands 5, yellow or pink, U-shaped, 0.2–0.3 × 0.4–0.5 mm; appendages greenish, white, or pink, forming narrow rim around gland, or ovate, oblong, or obovate and usually 2-fid, rarely rudimentary, 0.2–1.4 × 0.2–0.6 mm, entire. |
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Staminate flowers | 20–25. |
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Pistillate flowers | ovary glabrous, puberulent, strigillose, or pilose; styles 0.5–0.8 mm, 2-fid 1/3–1/2 length. |
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Capsules | oblate, 1.5–2.6 × 2.1–3.3 mm, glabrous or puberulent, strigillose, or pilose; columella 1.2–2.1 mm. |
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Seeds | brown to grayish black, narrowly ovoid, 3- or 4-angled in cross section, sometimes obscurely so, 1.3–1.9 × 1–1.4 mm, tuberculate, often with shallow depressions; caruncle absent. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at distal nodes or in weakly defined cymes or dichasia, dichasial bracts and distal stem leaves wholly green; peduncle 0.5–3.6 mm, strigillose. |
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2n | = 32. |
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Euphorbia polycarpa |
Euphorbia bilobata |
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Phenology | Flowering and fruiting spring–fall. | |
Habitat | Sandy and rocky soils on slopes and canyon bottoms in pine-juniper woodlands, oak woodlands, grasslands. | |
Elevation | 1400–2600 m. (4600–8500 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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AZ; NM; TX; Mexico (Chihuahua, Durango, Sonora)
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In Texas, Euphorbia bilobata is known only from Jeff Davis County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) |
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