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smallseed sandmat
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Herbs, annual, with taproot. |
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erect, usually branched near base, 6–22 cm, glabrous. |
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petiole absent; blade linear to slightly lanceolate or linear-oblanceolate, 5–18 × 0.5–2.5 mm, base linear attenuate, margins entire, apex rounded to obtuse or acute, surfaces glabrous; venation pinnate, midvein prominent. |
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infundibular, 0.8–1.1 × 0.6–0.9 mm, glabrous; glands 4, crescent-shaped; 0.2–0.4 × 0.5–0.6 mm; horns divergent, 0.5–0.7 mm. |
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5–10. |
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ovary glabrous; styles 0.3–0.5 mm, 2-fid. |
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ovoid-globose, 1.8–2.2 × 3–3.2 mm, slightly 3-lobed; cocci rounded, smooth, glabrous; columella 1.5–2 mm. |
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white to gray, ellipsoid, 1.4–1.7 × 1–1.3 mm, with deep, irregular to rounded, shallow to concave depressions over entire surface; caruncle reniform-ovate, depressed-conic, 0.5–0.7 × 0.7–1 mm. |
polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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(1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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reported two of these for the United States: var. hirtella and var. polycarpa. |
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hirtella, as the name suggests, was applied to hairy plants. |
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, it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts similar in shape to but slightly shorter and wider than distal leaves; dichasial bracts distinct, not imbricate, reniform-ovate to subdeltate-ovate or broadly ovate-lanceolate, base obliquely truncate to rounded, margins entire, apex obtuse to broadly acuminate; axillary cymose branches 0–3. |
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peduncle 0–0.5 mm. |
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AZ; CA; NV; nw Mexico
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TX |
Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia austrotexana occurs in stabilized sandy soil in the south Texas plains (M. H. Mayfield 2013). It is similar to E. longicruris but differs from that species in its often narrowly oblanceolate to linear leaves and its white to gray, ellipsoid seeds that are covered with minute, concave depressions. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Stems 10–22 cm; leaf blades linear-oblanceolate, apices rounded; dichasial bracts reniform-ovate to subdeltate-ovate, bases truncate; seeds 1.6–1.7 × 1.1–1.3 mm, surfaces with regular concave depressions; Atascosa, Bexar, Wilson counties. | var. austrotexana |
1. Stems 6–13 cm; leaf blades linear to slightly lanceolate, apices acute; dichasial bracts broadly ovate-lanceolate, bases rounded; seeds 1.4–1.5 × 1–1.1 mm, surfaces with irregular, not obviously concave depressions; Jim Hogg, Kenedy counties. | var. carrii |
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| FNA vol. 12, p. 283. |
FNA vol. 12, p. 298. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > subg. Esula |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
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| Chamaesyce polycarpa |
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| Bentham: Bot. Voy. Sulphur, 50. (1844) |
Mayfield: J. Bot. Res. Inst. Texas 7: 634, figs. 1, 2[row 3, left & center]. (2013) |
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