Euphorbia polycarpa |
Euphorbia antisyphilitica |
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smallseed sandmat |
candelilla, wax plant |
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Habit | Shrubs, with much-branched, fleshy rootstock. | |
Stems | erect, few branched, 25–50(–100) cm, glabrous or puberulent, pencil-like, in age covered with flaky, exfoliating layer of wax. |
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Leaves | alternate usually caducous, sometimes persisting; stipules 0.4–0.5 mm; petiole absent; blade ovate to deltate-subulate, 2.5–4 × 1 mm, thick, fleshy, base usually rounded and swollen, rarely cuneate, margins entire, apex acute, surfaces puberulent, adaxial sometimes canescent; venation inconspicuous. |
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Involucre | campanulate, 1.6–2.2 × 1.6–1.9 mm, puberulent to canescent; glands 5, pinkish, narrowly oblong to reniform, 0.3–0.4 × 0.8–1 mm; appendages white to pink, ovate, oblong, or transversely oblong, 1.3–2.5 × 1.4–2.5 mm, usually erose, rarely entire. |
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Staminate flowers | 50–70. |
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Pistillate flowers | ovary glabrous; styles 0.9–1.1 mm, 2-fid nearly entire length. |
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Capsules | oblong to ovoid, 3.9–4.2 × 3.6–3.9 mm, glabrous; columella 3.1–3.3 mm. |
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Seeds | whitish gray, narrowly ovoid, 2.4–3.1 × 1.4–1.6 mm, irregularly rugose-tuberculate; caruncle crescent-shaped, 0.3–0.6 × 0.6–0.8 mm. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | in axillary congested cymes, near branch tips or solitary at distal nodes; peduncle 0–1 mm, lanulose. |
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Euphorbia polycarpa |
Euphorbia antisyphilitica |
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Phenology | Flowering and fruiting year-round in response to sufficient rainfall. | |
Habitat | Desert scrub, frequently on limestone substrates. | |
Elevation | 100–1200 m. (300–3900 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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NM; TX; Mexico
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia antisyphilitica is the only pencil-stemmed species of Euphorbia occurring in the flora area. The species is characteristic of the Chihuahuan Desert scrub of Mexico from Chihuahua and Coahuila south to Hidalgo and Querétaro, and barely enters into the United States in southern New Mexico (Doña Ana and Lincoln counties) and southwest (Brewster, Hudspeth, Presidio, and Terrell counties) and south (Starr and Webb counties) Texas. The stems are covered in a conspicuous coat of exfoliating wax, and the plants historically have been harvested for this product, although the practice is much less prevalent now. The specific epithet refers to its traditional medicinal use in treating sexually transmitted infections. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 243. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Zuccarini: Flora 15(2, Beibl.): 58. (1832) |
Web links |