Euphorbia polycarpa |
Euphorbia acuta |
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smallseed sandmat |
point sandmat |
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Habit | Herbs, perennial, with strongly thickened rootstock. | |
Stems | ascending to erect, 5–30 cm, uniformly and densely canescent or sericeous. |
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Leaves | opposite; stipules deciduous, sometimes appearing absent, distinct, brown, linear-subulate, thin, 0.3–0.8 mm, canescent; petiole 0.4–1.2 mm, moderately to densely canescent; blade ovate to lanceolate, 6–20 × 3–8 mm, base subsymmetric, rounded to cuneate, margins entire, strongly involute, apex long-acuminate, spinulose, abaxial surface canescent to densely sericeous, adaxial surface glabrous or sparsely canescent; 3-veined from base but only midvein conspicuous. |
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Involucre | turbinate to urceolate, 2–2.6 × 1.7–2.5 mm, villous to lanate; glands 4, yellow-green to orange or red, slightly concave, oblong-elliptic, 0.2–0.4 × 0.6–1.5 mm; appendages white, flabellate, 1.1–2.1 × 0.2–0.6 mm, distal margin shallowly and irregularly toothed. |
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Staminate flowers | 20–25. |
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Pistillate flowers | ovary strigose, pubescent to villous; styles 0.6–0.9 mm, 2-fid 1/2 length. |
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Capsules | subglobose to broadly ovoid, 2.8–3.7 mm diam., strigose, pubescent to villous; columella 2.3–3 mm. |
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Seeds | white, ovoid, 4-angled in cross section, 2.2–2.6 × 1.1–1.4 mm, smooth to finely reticulate. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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Cyathia | solitary at distal nodes; peduncle 1.3–3.2 mm. |
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2n | = 28, 48, 56. |
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Euphorbia polycarpa |
Euphorbia acuta |
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Phenology | Flowering and fruiting spring–fall. | |
Habitat | Desert scrub, grasslands, oak-juniper savannas, limestone, rocky, sandy, or clay soils. | |
Elevation | 400–1900 m. (1300–6200 ft.) | |
Distribution |
AZ; CA; NV; nw Mexico
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NM; TX; Mexico (Chihuahua, Coahuila) |
Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia acuta is easily distinguished in the field by its relatively large, strongly involute, hairy, and acutely pointed leaves. The name Euphorbia acuta Engelmann has been proposed for conservation against the earlier name E. acuta Bellardi ex Colla (P. E. Berry et al. 2011). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 283. | FNA vol. 12, p. 259. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce polycarpa | Chamaesyce acuta, E. acuta var. stenophylla, E. georgei |
Name authority | Bentham: Bot. Voy. Sulphur, 50. (1844) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) — name proposed for conservation |
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