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perennial sandmat, Terlingua spurge

Mendez's sandmat

Habit Herbs, perennial, with strongly thickened, woody rootstock. Herbs, annual or perennial, with slender taproot.
Stems

erect, 7–45 cm, glabrous.

prostrate, often mat-forming, 8–35 cm, usually villous along margins, lower surface glabrous, upper surface usually strigillose to puberulent, rarely glabrous or glabrate.

Leaves

opposite;

stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous;

petiole 0.8–2 mm, glabrous;

blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous.

opposite;

stipules usually distinct or connate basally, rarely completely connate, deltate, laciniate, glabrous or pilose (lower side), forming narrow deltate scale, sometimes apically 2-fid or laciniate, glabrous (upper side), 0.4–1.9 mm;

petiole 0.3–1.2 mm, glabrous, pilose or villous;

blade oblong to obovate, 4–12 × 2–7 mm, base asymmetric, one side attenuate, cuneate or rounded, other rounded or cordate, margins serrulate at least distally, apex obtuse, surfaces glabrous or sparsely sericeous, pilose or villous; 3-veined at base.

Involucre

broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous;

glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm;

appendages absent.

campanulate or obconic, 0.8–1 × 0.7–0.8 mm, usually glabrous, rarely sparsely pilose toward apex;

glands 4, pink, reniform, oblong or elliptic, 0.1 × 0.2–0.3 mm;

appendages absent or white to pink, oblong, flabellate or forming narrow rim around edge of gland, 0.1–0.3 × 0.3–0.6 mm, distal margin usually entire, sometimes lobed.

Staminate flowers

35–45.

6–15.

Pistillate flowers

ovary glabrous;

styles 0.7–0.9 mm, 2-fid nearly entire length.

ovary usually pilose or villous with hairs concentrated along keels, rarely glabrous;

styles 0.2–0.3 mm, 2-fid 1/2 length.

Capsules

subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous;

columella 2.2–2.7 mm.

ovoid, 1.2–1.6 × 1.2–1.4 mm, usually pilose or villous with hairs concentrated along keels, often glabrous in between, very rarely completely glabrous;

columella 1–1.4 mm.

Seeds

white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled.

reddish brown to orange or gray-pink, narrowly ovoid, 4-angled in cross section, 0.9–1.2 × 0.5–0.6 mm, almost smooth or with 5–7 faint transverse ridges that do not pass through abaxial keel.

Cyathia

solitary at distal nodes or at nodes of short, axillary branches;

peduncle 1.8–3 mm.

solitary at nodes or on short, congested axillary branches;

peduncle 0.9–2.5 mm.

Euphorbia perennans

Euphorbia mendezii

Phenology Flowering and fruiting spring–fall. Flowering and fruiting year-round.
Habitat Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. Disturbed areas.
Elevation 900–1200 m. (3000–3900 ft.) 0–10 m. (0–0 ft.)
Distribution
from FNA
TX; Mexico (Chihuahua)
[BONAP county map]
from FNA
FL; Mexico; Central America; West Indies; n South America [Introduced in North America]
[BONAP county map]
Discussion

Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011).

Euphorbia perennans is known in the flora area only from Brewster County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia mendezii is a common weed distributed widely throughout Mexico and Central America. Within the flora area the species is known only from southern Florida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 282. FNA vol. 12, p. 277.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce perennans Chamaesyce mendezii
Name authority (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) Boissier: Cent. Euphorb., 15. (1860)
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