Euphorbia perennans |
Euphorbia inundata |
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perennial sandmat, Terlingua spurge |
Florida pineland spurge |
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Habit | Herbs, perennial, with strongly thickened, woody rootstock. | Herbs, perennial, with thickened rootstock. | ||||
Stems | erect, 7–45 cm, glabrous. |
erect or ascending, 15–40 cm. |
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Leaves | opposite; stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous; petiole 0.8–2 mm, glabrous; blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous. |
petiole absent or indistinct, blade linear to linear-elliptic or lanceolate, (25–)30–60(–115) × 1.5–14(–15) mm, chartaceous, base attenuate, apex acuminate, acute, or rounded and mucronate; only midvein evident. |
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Involucre | broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous; glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm; appendages absent. |
obconic or campanulate, 1.3–3.2 × 1.6–3.6 mm, lobes ovate, oblong, or nearly rectangular, 0.6–1 mm, laciniate-ciliate; glands red to greenish, oblong or nearly circular, 0.8–1 × 1.1–2 mm, distal margins crenulate-erose, undulate, or entire. |
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Staminate flowers | 35–45. |
20–25. |
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Pistillate flowers | ovary glabrous; styles 0.7–0.9 mm, 2-fid nearly entire length. |
gynophore exserted 2.6–6.1 mm, calyxlike lobes triangular to subulate, 0.6–1.4 mm; styles connate 1/8 length, 1.3–2.2 mm. |
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Capsules | subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous; columella 2.2–2.7 mm. |
oblate-ovoid, 5.1–6.1 × 6.9–8.6 mm, 3-lobed; columella 3.9–4.7 mm. |
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Seeds | white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled. |
brown to blackish, ovoid-globose, circular or faintly 3- or 4-angled in cross section, 2.9–3.4 × 2.7–3 mm, smooth, base flattened or rounded, apex rounded or with inconspicuous blunt point. |
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Cyathia | solitary at distal nodes or at nodes of short, axillary branches; peduncle 1.8–3 mm. |
peduncle 6–24 mm (often exceeding dichasial bracts). |
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Cyathial | arrangement: terminal pleiochasial branches 2–3, 5–18 cm, 3–8 times 2-branched; pleiochasial bracts linear-lanceolate or narrowly ovate, 25–49 × 5–9 mm, margins entire, apex acute or acuminate; dichasial bracts ovate or lanceolate, 6–21 × 2–6 mm, margins entire, apex acute or acuminate; axillary cymose branches 1–5. |
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Euphorbia perennans |
Euphorbia inundata |
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Phenology | Flowering and fruiting spring–fall. | |||||
Habitat | Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. | |||||
Elevation | 900–1200 m. (3000–3900 ft.) | |||||
Distribution |
TX; Mexico (Chihuahua) |
AL; FL; GA; MS |
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Discussion | Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011). Euphorbia perennans is known in the flora area only from Brewster County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia inundata has a disjunct distribution in the southeastern United States. Allopatric, narrow-leaved populations from the west-central Florida peninsula are segregated as var. garrettii (E. L. Bridges and S. L. Orzell 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 282. | FNA vol. 12, p. 314. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce perennans | Galarhoeus inundatus | ||||
Name authority | (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) | Torrey ex Chapman: Fl. South. U.S., 402. (1860) | ||||
Web links |