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perennial sandmat, Terlingua spurge

dwarf spurge, small spurge

Habit Herbs, perennial, with strongly thickened, woody rootstock. Herbs, annual, with taproot.
Stems

erect, 7–45 cm, glabrous.

erect, unbranched or branched, 3–30 cm, glabrous.

Leaves

opposite;

stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous;

petiole 0.8–2 mm, glabrous;

blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous.

petiole 0–0.5 mm;

blade linear, linear-oblong, or linear-spatulate, 2–30 × 1–5 mm, base cuneate or truncate, margins entire, apex acute, obtuse, or emarginate, surfaces glabrous;

venation inconspicuous, only midvein prominent.

Involucre

broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous;

glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm;

appendages absent.

cupulate, 0.3–0.5 × 0.6–0.8 mm, glabrous;

glands 4, elliptic to crescent-shaped, 0.2–0.4 × 0.3–0.6 mm;

horns divergent, 0.2–0.5 mm.

Staminate flowers

35–45.

5–8.

Pistillate flowers

ovary glabrous;

styles 0.7–0.9 mm, 2-fid nearly entire length.

ovary glabrous;

styles 0.5–0.7 mm, 2-fid.

Capsules

subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous;

columella 2.2–2.7 mm.

subglobose, 1–1.8 × 1.3–2 mm, slightly lobed;

cocci rounded, smooth, puncticulate toward abaxial line, glabrous;

columella 1.1–1.4 mm.

Seeds

white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled.

blackish to grayish, 4-angled-ovoid, 1–1.5 × 0.5–0.7 mm, white, tuberculate;

caruncle conic or subconic, 0.1–0.3 × 0.1–0.3 mm.

Cyathia

solitary at distal nodes or at nodes of short, axillary branches;

peduncle 1.8–3 mm.

peduncle 0–2 mm.

Cyathial

arrangement: terminal pleiochasial branches 3–5, each 1–3 times 2-branched;

pleiochasial bracts similar in shape and size to or sometimes slightly longer and wider than distal leaves;

dichasial bracts distinct, linear, or linear-lanceolate, or lanceolate-ovate, base rounded to subcordate, margins entire, apex acute;

axillary cymose branches 0–5.

Euphorbia perennans

Euphorbia exigua

Phenology Flowering and fruiting spring–fall. Flowering and fruiting summer–fall.
Habitat Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. Edges of gardens, roadsides, waste places.
Elevation 900–1200 m. (3000–3900 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
TX; Mexico (Chihuahua)
[BONAP county map]
from FNA
CA; NY; WV; BC; NS; ON; Europe; w Asia; n Africa; Atlantic Islands (Macaronesia) [Introduced in North America]
[BONAP county map]
Discussion

Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011).

Euphorbia perennans is known in the flora area only from Brewster County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia exigua can be easily distinguished from other annual species of the genus in the flora area by its tuberculate seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 282. FNA vol. 12, p. 302.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce perennans Tithymalus exiguus
Name authority (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) Linnaeus: Sp. Pl. 1: 456. (1753)
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