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perennial sandmat, Terlingua spurge
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summer spurge
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| Herbs, perennial, with strongly thickened, woody rootstock. |
Herbs, perennial, with spreading rootstock. |
erect, 7–45 cm, glabrous. |
erect or ascending, unbranched, solitary or few, previous year's dead stems not persistent, 45–70 cm, usually densely puberulent to sericeous, rarely glabrous. |
opposite; stipules distinct, linear-filiform in (1–)2(–3) segments, 0.3–0.4 mm, glabrous; petiole 0.8–2 mm, glabrous; blade ovate or orbiculate-deltate to reniform-deltate, 5–17 × 4–16 mm midstem leaves largest, base symmetric, cuneate, rounded to cordate, margins entire, apex acute to rounded, surfaces glabrous, often glaucous; 3-veined from base, only midvein conspicuous. |
alternate; stipules to 0.1 mm; petiole (0–)1–2 mm (or absent), densely puberulent; blade usually linear, rarely ovate, 25–55 × 1.5–4 mm, base cuneate, margins entire, revolute, apex rounded, abaxial surface glabrous or puberulent to sericeous, adaxial surface glabrous; venation often obscure on smaller leaves, midvein conspicuous. |
broadly-campanulate to hemispheric, 1.7–2.2 × 1.5–2.7 mm, glabrous; glands 4, green to yellow-green, elliptic to oblong, folded longitudinally, 0.3–0.5 × 0.7–1.4 mm; appendages absent. |
campanulate, 1.2–1.4 × 1.2–2 mm, sparsely to densely puberulent; glands 5, green, reniform, 0.2–0.3 × 0.5–0.6 mm; appendages white, orbiculate to oblong, (0.5–)1–1.7 × 1–1.5 mm, entire. |
35–45. |
20–25. |
ovary glabrous; styles 0.7–0.9 mm, 2-fid nearly entire length. |
ovary glabrous or sparsely strigose; styles 0.5–1.1 mm, 2-fid at apex to 1/2 length. |
subglobose to broadly ovoid, 2.8–3.3 × 2.8–3.4 mm, glabrous; columella 2.2–2.7 mm. |
globose, 1.8–3 × 2.5–4.8 mm, glabrous or sparsely strigose; columella 2.3–2.5 mm. |
white to light brown, ovoid, 3–4-angled in cross section, 2–2.4 × 1–1.2 mm, smooth to faintly transverse-wrinkled. |
light gray, ovoid, 2 × 1.2–1.3 mm, smooth or with few, very shallow depressions; caruncle absent. |
solitary at distal nodes or at nodes of short, axillary branches; peduncle 1.8–3 mm. |
in terminal pleiochasia; peduncle 5–15 mm, filiform, glabrous or very sparsely puberulent to sericeous. |
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| Flowering and fruiting spring–fall. |
Flowering and fruiting late spring–fall. |
| Desert scrub, on cretaceous and gypseous clay, limestone hills and flats. |
Sand hills, pine savannas, woodland borders, open fields with sandy soils. |
| 900–1200 m. (3000–3900 ft.) |
0–150 m. (0–500 ft.) |
|
TX; Mexico (Chihuahua) |
AL; FL; GA; LA; MS; TX
|
Euphorbia perennans is a distinctive species with an erect habit and relatively large, firm, deltate midstem leaves. Phylogenetic data place E. perennans in a clade of primarily Chihuahuan Desert annual and perennial species (for example, E. chaetocalyx, E. fendleri, E. golondrina, E. simulans, E. spurca, and E. theriaca; Y. Yang and P. E. Berry 2011). Euphorbia perennans is known in the flora area only from Brewster County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
M. J. Huft (1979) remarked that Euphorbia discoidalis is uncommon west of Alabama and referred many narrow-leaved specimens from Louisiana and Texas to E. corollata. K. R. Park (1998) included them in an expanded E. discoidalis, and that is followed here. The western populations can be distinguished from E. corollata by their shorter involucral gland appendages and revolute leaf margins. Further study of these western populations is warranted. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 282. |
FNA vol. 12, p. 245. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| Chamaesyce perennans |
Tithymalopsis discoidalis |
| (Shinners) Warnock & M. C. Johnston: SouthW. Naturalist 5: 170. (1960) |
Chapman: Fl. South. U.S., 401. (1860) |
| |
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