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Carrizo Mountain sandmat

sand-dune sandmat or spurge

Habit Herbs, annual, with taproot.
Stems

prostrate, spreading and lanky or occasionally mat-forming, ± succulent, 70–150 cm, glabrous.

Leaves

opposite;

stipules usually distinct, occasionally connate basally (primarily at distal nodes), usually divided into 2–5 subulate to subulate-filiform segments, occasionally forming narrow deltate segments (primarily at distal nodes), 0.8–1.8 mm, glabrous;

petiole 3.1–6.3 mm, glabrous;

blade ovate to elliptic-oblong, 5–26 × 4–12 mm, base subsymmetric to symmetric, rounded to cuneate, margins entire, apex usually mucronate, rarely acute or obtuse, surfaces glabrous; pinnately veined.

Involucre

campanulate, 1.1–1.8 × 1.5–2.5 mm, glabrous;

glands 4, yellowish, sessile or short stipitate, circular to oblong, 0.5–0.7 × 0.5–0.8 mm;

appendages white to yellowish, ovate to oblong, occasionally rudimentary, (0–)0.8–1.2 × 0.8–1.5 mm, distal margin entire.

Staminate flowers

15–25.

Pistillate flowers

ovary glabrous;

styles 0.7–1 mm, 2-fid 1/2 length.

Capsules

ovoid, 4.7–5.5(–6) × 3.6–5.1 mm, glabrous;

columella 4.3–5.1 mm.

Seeds

grayish white to reddish brown mottled, bottle-shaped, strongly dorsiventrally compressed and weakly 3-angled in cross section, (2.8–)4.1–5.2 × 1.2–2(–3.4) mm, smooth; carunclelike structure linear, 0.4–0.5 × 0.1–0.2 mm.

Euphorbia

pediculifera can be variable in terms of leaf shape and the extent of pubescence, but the species is readily identified by the short, appressed hairs on the leaves and the distinctive seed that is encircled by a series of round, transverse ridges.

Cyathia

solitary at distal nodes;

peduncle 1.4–3.6 mm.

Euphorbia pediculifera

Euphorbia carunculata

Phenology Flowering and fruiting summer–fall.
Habitat Sand dunes.
Elevation 400–1300 m. (1300–4300 ft.)
Distribution
from FNA
AZ; CA; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
KS; NM; OK; TX; Mexico (Chihuahua)
[BONAP county map]
Discussion

Varieties 2 (1 in the flora).

Euphorbia pediculifera var. linearifolia S. Watson is found only near Guaymas in southwestern Sonora, Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia carunculata has a highly localized and scattered distribution. The species is restricted to sand dunes and known from only a handful of localities throughout its relatively wide range. The seeds are unique in being bottle-shaped and strongly laterally compressed, and unlike other members of sect. Anisophyllum, there is a minute, linear, carunclelike protuberance at the hilum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 281. FNA vol. 12, p. 262.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. pediculifera var. pediculifera
Synonyms Chamaesyce pediculifera Chamaesyce carunculata
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) Waterfall: Rhodora 50: 63. (1948)
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