Euphorbia pediculifera |
Euphorbia brachycera |
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Carrizo Mountain sandmat |
horned spurge, shorthorn spurge |
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Habit | Herbs, perennial, with thick rootstock. | |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
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Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
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Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
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Staminate flowers | 10–15. |
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Pistillate flower | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
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Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
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Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
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Euphorbia | pediculifera can be variable in terms of leaf shape and the extent of pubescence, but the species is readily identified by the short, appressed hairs on the leaves and the distinctive seed that is encircled by a series of round, transverse ridges. |
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Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
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Cyathia | peduncle 0.4–1 mm. |
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2n | = 28. |
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Euphorbia pediculifera |
Euphorbia brachycera |
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Phenology | Flowering and fruiting spring–summer. | |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | |
Elevation | 1200–3200 m. (3900–10500 ft.) | |
Distribution |
AZ; CA; nw Mexico
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AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
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Discussion | Varieties 2 (1 in the flora). Euphorbia pediculifera var. linearifolia S. Watson is found only near Guaymas in southwestern Sonora, Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 281. | FNA vol. 12, p. 300. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Chamaesyce pediculifera | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) |
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