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dune spurge, Parry's sandmat, Parry's spurge

Roemer's spurge

Habit Herbs, annual, with taproot. Herbs, annual, with taproot.
Stems

usually prostrate, rarely ascending-erect, 5–70(–85) cm, glabrous.

erect, occasionally decumbent at base, branched, 15–30 cm, glabrous.

Leaves

opposite;

stipules distinct, linear-subulate, usually lacerate and divided into 2 or more slender segments, rarely entire, 0.6–1.4 mm, glabrous;

petiole 1–2.5 mm, glabrous;

blade linear to narrowly oblong, (5–)10–25(–30) × 2–5 mm, base usually symmetric, sometimes slightly asymmetric, attenuate, margins entire, occasionally ± revolute, apex acute to obtuse, mucronulate, surfaces glabrous;

only midvein conspicuous.

petiole 2–5 mm;

blade oblanceolate to obovate, 5–20 × 5–10 mm (larger leaves in distal portion of stem), base cuneate to attenuate, margins entire, apex rounded, surfaces glabrous;

venation pinnate.

Involucre

broadly cupuliform-campanulate, 1.2–1.7 × 1.4–1.8 mm, glabrous;

glands 4, reddish pink to greenish yellow, deeply concave, elliptic to oblong, 0.2–0.3 × 0.3–0.5 mm;

appendages white, elliptic to oblong, usually forming narrow margin around gland, sometimes rudimentary, 0.2–0.6 × 0.3–0.7(–1.1) mm, distal margin entire.

campanulate, 1.5–2 × 1–1.6 mm, glabrous;

glands 4, elliptic to trapezoidal, 0.4–0.6 × 0.7–1.2 mm;

horns slightly convergent, 0.1–0.3 mm.

Staminate flowers

40–55.

10–12.

Pistillate flowers

ovary glabrous;

styles 0.5–0.7 mm, 2-fid nearly entire length.

ovary glabrous;

styles 0.6–1 mm, 2-fid.

Capsules

ovoid-globose, 2–2.3 × 1.5–2.5 mm, glabrous;

columella 1.4–2 mm.

subglobose, 2–2.5 × 2.4–3 mm, slightly lobed;

cocci rounded, smooth, glabrous;

columella 1.5–1.8 mm.

Seeds

mottled brown and white because of irregularly loose and tight outer covering, broadly ovoid, rounded-angular in cross section, 1.4–1.8 × 0.8–1 mm, smooth or only inconspicuously roughened.

brown, oblong-ovoid, 1.6–1.8 × 1.4–1.5 mm, with scattered, deep and broad pits;

caruncle conic, reniform, 0.3–0.4 × 0.4–0.6 mm.

Cyathia

solitary or in small clusters on short axillary branches at distal nodes;

peduncle 1–5 mm.

peduncle 0–0.9 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, 1–4 times 2-branched;

pleiochasial bracts ovate to oblong, similar in size to distal leaves;

dichasial bracts connate 1/2 length, reniform to semicircular, base truncate to perfoliate, margins entire, apex rounded to obtuse;

axillary cymose branches 0–3.

Euphorbia parryi

Euphorbia roemeriana

Phenology Flowering and fruiting spring–fall. Flowering and fruiting spring.
Habitat Sand dunes, other sandy habitats. Rich calcareous soils, creek canyons.
Elevation 200–2200 m. (700–7200 ft.) 100–300 m. (300–1000 ft.)
Distribution
from FNA
AZ; CA; CO; NM; NV; TX; UT; Mexico (Chihuahua)
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Euphorbia parryi is similar to E. missurica, differing only by the generally narrow involucral gland appendages and prostrate habit in E. parryi as opposed to the conspicuous involucral gland appendages and ascending-erect habit in E. missurica. Euphorbia parryi has sometimes been considered the western race of E. missurica (D. S. Correll and M. C. Johnston 1970).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

D. S. Correll and M. C. Johnston (1970) suggested that Euphorbia roemeriana, which is restricted to the eastern part of the Edwards Plateau, is the southern counterpart of E. commutata, to which it is morphologically very similar. However, molecular phylogenetic analyses indicate that it is most closely related to E. austrotexana and E. longicruris (J. A. Peirson et al. 2014). Euphorbia roemeriana can be distinguished from E. commutata and several of the other small, annual species of subg. Esula by its consistently connate dichasial bracts.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 281. FNA vol. 12, p. 308.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce longeramosa, C. parryi, E. longeramosa Tithymalus roemerianus
Name authority Engelmann: Amer. Naturalist 9: 350. (1875) Scheele: Linnaea 22: 151. (1849)
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