Euphorbia parryi |
Euphorbia revoluta |
|
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dune spurge, Parry's sandmat, Parry's spurge |
revolute spurge, threadstem sandmat, threadstem spurge |
|
Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | usually prostrate, rarely ascending-erect, 5–70(–85) cm, glabrous. |
erect, 5–25 cm, glabrous. |
Leaves | opposite; stipules distinct, linear-subulate, usually lacerate and divided into 2 or more slender segments, rarely entire, 0.6–1.4 mm, glabrous; petiole 1–2.5 mm, glabrous; blade linear to narrowly oblong, (5–)10–25(–30) × 2–5 mm, base usually symmetric, sometimes slightly asymmetric, attenuate, margins entire, occasionally ± revolute, apex acute to obtuse, mucronulate, surfaces glabrous; only midvein conspicuous. |
opposite; stipules distinct, subulate-filiform, entire, 0.5–0.9 mm, glabrous; petiole 0.6–1.1 mm, glabrous; blade linear, 6–27 × 0.6–1.2 mm, base symmetric, attenuate, margins entire, revolute, apex acute, sometimes mucronate, surfaces glabrous; only midvein conspicuous. |
Involucre | broadly cupuliform-campanulate, 1.2–1.7 × 1.4–1.8 mm, glabrous; glands 4, reddish pink to greenish yellow, deeply concave, elliptic to oblong, 0.2–0.3 × 0.3–0.5 mm; appendages white, elliptic to oblong, usually forming narrow margin around gland, sometimes rudimentary, 0.2–0.6 × 0.3–0.7(–1.1) mm, distal margin entire. |
obconic to campanulate, 0.7–0.9 × 0.5–0.7 mm, glabrous; glands 4, pink to dark purple, nearly circular to oblong or reniform, 0.1 × 0.1–0.2 mm; appendages white, oblong, nearly circular, ovate, deltate, or forming thin margin around gland, rarely absent, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin entire. |
Staminate flowers | 40–55. |
5–10. |
Pistillate flowers | ovary glabrous; styles 0.5–0.7 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.2–0.3 mm, unbranched. |
Capsules | ovoid-globose, 2–2.3 × 1.5–2.5 mm, glabrous; columella 1.4–2 mm. |
ovoid to subglobose, 1.5–1.8 × 1.6–1.8 mm, glabrous; columella 1.1–1.5 mm. |
Seeds | mottled brown and white because of irregularly loose and tight outer covering, broadly ovoid, rounded-angular in cross section, 1.4–1.8 × 0.8–1 mm, smooth or only inconspicuously roughened. |
whitish, brick red, light gray, or light brown, narrowly to broadly ovoid, 4-angled in cross section, 0.9–1.4 × 0.7–1 mm, nearly smooth, rugulose, with faint transverse ridges, or with 2–3 well-defined transverse ridges separated by shallow depressions. |
Cyathia | solitary or in small clusters on short axillary branches at distal nodes; peduncle 1–5 mm. |
solitary at distal nodes; peduncles absent or to 1.5 mm. |
Euphorbia parryi |
Euphorbia revoluta |
|
Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting summer–late fall. |
Habitat | Sand dunes, other sandy habitats. | Desert scrub, sagebrush scrub, juniper woodlands, Joshua tree-pinyon pine woodlands, oak woodlands, grasslands, chaparral, pine-oak forests. |
Elevation | 200–2200 m. (700–7200 ft.) | 600–2500 m. (2000–8200 ft.) |
Distribution |
AZ; CA; CO; NM; NV; TX; UT; Mexico (Chihuahua)
|
AZ; CA; CO; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Sonora, Zacatecas)
|
Discussion | Euphorbia parryi is similar to E. missurica, differing only by the generally narrow involucral gland appendages and prostrate habit in E. parryi as opposed to the conspicuous involucral gland appendages and ascending-erect habit in E. missurica. Euphorbia parryi has sometimes been considered the western race of E. missurica (D. S. Correll and M. C. Johnston 1970). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia revoluta is a distinctive, easily recognizable species by virtue of the combination of unbranched styles and linear leaves with symmetric bases. The species ranges through northern Mexico into the southwestern United States and is composed of three well-marked, geographically distinct races that can be distinguished by their seeds. The first of these races occurs primarily in pine-oak forest of the Sierra Madre Occidental of Mexico (Chihuahua and Sonora) and barely enters the flora area in the Huachuca Mountains of southeastern Arizona; the seeds are brick red and nearly smooth. The second race is widespread in the Chihuahuan Desert from northern Mexico to southeastern New Mexico and southwestern Texas; its seeds are whitish to light gray and possess two or three well-defined transverse ridges separated by shallow depressions. The third race corresponds to the type collection and is characterized by grayish white to light brown seeds that are rugulose or with faint transverse ridges; it occurs in northwestern Mexico (Baja California and Sonora) and throughout the arid southwestern United States. It is probable that further study will justify the taxonomic segregation of these races as either distinct species or subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 281. | FNA vol. 12, p. 286. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce longeramosa, C. parryi, E. longeramosa | Chamaesyce revoluta |
Name authority | Engelmann: Amer. Naturalist 9: 350. (1875) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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