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dune spurge, Parry's sandmat, Parry's spurge
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Heller's spurge
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| Herbs, annual, with taproot. |
Herbs, annual, with taproot. |
usually prostrate, rarely ascending-erect, 5–70(–85) cm, glabrous. |
ascending, branched proximally, 15–30 cm, glabrous. |
opposite; stipules distinct, linear-subulate, usually lacerate and divided into 2 or more slender segments, rarely entire, 0.6–1.4 mm, glabrous; petiole 1–2.5 mm, glabrous; blade linear to narrowly oblong, (5–)10–25(–30) × 2–5 mm, base usually symmetric, sometimes slightly asymmetric, attenuate, margins entire, occasionally ± revolute, apex acute to obtuse, mucronulate, surfaces glabrous; only midvein conspicuous. |
petiole usually 1–3 mm, absent distally; blade spatulate, 6–15 × 3–5 mm, base broadly attenuate, margins entire, apex usually obtuse to rounded, sometimes retuse, surfaces glabrous; venation pinnate, midvein prominent. |
broadly cupuliform-campanulate, 1.2–1.7 × 1.4–1.8 mm, glabrous; glands 4, reddish pink to greenish yellow, deeply concave, elliptic to oblong, 0.2–0.3 × 0.3–0.5 mm; appendages white, elliptic to oblong, usually forming narrow margin around gland, sometimes rudimentary, 0.2–0.6 × 0.3–0.7(–1.1) mm, distal margin entire. |
campanulate, 0.9–1.2 × 0.7–1 mm, glabrous; glands 4, elliptic, 0.1–0.2 × 0.3–0.4 mm; horns slightly divergent, 0.1–0.2 mm. |
40–55. |
8–10. |
ovary glabrous; styles 0.5–0.7 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.4–0.5 mm, 2-fid. |
ovoid-globose, 2–2.3 × 1.5–2.5 mm, glabrous; columella 1.4–2 mm. |
depressed-globose, 2.1–2.5 × 2.5–3 mm, 3-lobed; cocci flattened, smooth, glabrous; columella 1.2–1.7 mm. |
mottled brown and white because of irregularly loose and tight outer covering, broadly ovoid, rounded-angular in cross section, 1.4–1.8 × 0.8–1 mm, smooth or only inconspicuously roughened. |
white to light gray, ovoid, 1.4–1.6 × 1–1.2 mm, smooth; caruncle 2-lobed, thin, 0.4 × 0.7 mm. |
solitary or in small clusters on short axillary branches at distal nodes; peduncle 1–5 mm. |
peduncle 0.2–0.4 mm. |
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arrangement: terminal pleiochasial branches 3, each 1–5 times 2-branched; pleiochasial bracts oblong, similar in size to distal leaves; dichasial bracts distinct, orbiculate-ovate to nearly reniform, subpandurate, base broadly cuneate to truncate, margins entire, apex obtuse, mucronate; axillary cymose branches 0–5. |
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| Flowering and fruiting spring–fall. |
Flowering and fruiting late winter–spring. |
| Sand dunes, other sandy habitats. |
Forests, stream banks, roadsides, shaded areas with sandy, calcareous soils. |
| 200–2200 m. (700–7200 ft.) |
0–50 m. (0–200 ft.) |
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AZ; CA; CO; NM; NV; TX; UT; Mexico (Chihuahua)
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LA; TX; Mexico (Nuevo León) |
Euphorbia parryi is similar to E. missurica, differing only by the generally narrow involucral gland appendages and prostrate habit in E. parryi as opposed to the conspicuous involucral gland appendages and ascending-erect habit in E. missurica. Euphorbia parryi has sometimes been considered the western race of E. missurica (D. S. Correll and M. C. Johnston 1970). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The smooth, white to light gray seeds of Euphorbia helleri easily distinguish it from other annual members of subg. Esula in North America. Collections of E. helleri have been made near Brownsville, Texas, and thus it is possible that the species occurs in northern Tamaulipas, Mexico. The Louisiana record (Webster Parish) likely represents introduced plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 281. |
FNA vol. 12, p. 303. |
| Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Euphorbiaceae > Euphorbia > subg. Esula |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| Chamaesyce longeramosa, C. parryi, E. longeramosa |
Tithymalus helleri |
| Engelmann: Amer. Naturalist 9: 350. (1875) |
Millspaugh: Bot. Gaz. 26: 268, fig. [p. 270]. (1898) |
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