Euphorbia parryi |
Euphorbia exstipulata |
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dune spurge, Parry's sandmat, Parry's spurge |
square-seed spurge, squareseed or Clark Mountain spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | usually prostrate, rarely ascending-erect, 5–70(–85) cm, glabrous. |
erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
Leaves | opposite; stipules distinct, linear-subulate, usually lacerate and divided into 2 or more slender segments, rarely entire, 0.6–1.4 mm, glabrous; petiole 1–2.5 mm, glabrous; blade linear to narrowly oblong, (5–)10–25(–30) × 2–5 mm, base usually symmetric, sometimes slightly asymmetric, attenuate, margins entire, occasionally ± revolute, apex acute to obtuse, mucronulate, surfaces glabrous; only midvein conspicuous. |
opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
Involucre | broadly cupuliform-campanulate, 1.2–1.7 × 1.4–1.8 mm, glabrous; glands 4, reddish pink to greenish yellow, deeply concave, elliptic to oblong, 0.2–0.3 × 0.3–0.5 mm; appendages white, elliptic to oblong, usually forming narrow margin around gland, sometimes rudimentary, 0.2–0.6 × 0.3–0.7(–1.1) mm, distal margin entire. |
turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
Staminate flowers | 40–55. |
10–12. |
Pistillate flowers | ovary glabrous; styles 0.5–0.7 mm, 2-fid nearly entire length. |
ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
Capsules | ovoid-globose, 2–2.3 × 1.5–2.5 mm, glabrous; columella 1.4–2 mm. |
broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
Seeds | mottled brown and white because of irregularly loose and tight outer covering, broadly ovoid, rounded-angular in cross section, 1.4–1.8 × 0.8–1 mm, smooth or only inconspicuously roughened. |
white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
Cyathia | solitary or in small clusters on short axillary branches at distal nodes; peduncle 1–5 mm. |
peduncle 1–1.9 mm. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
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Euphorbia parryi |
Euphorbia exstipulata |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting summer–fall. |
Habitat | Sand dunes, other sandy habitats. | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. |
Elevation | 200–2200 m. (700–7200 ft.) | 800–2000 m. (2600–6600 ft.) |
Distribution |
AZ; CA; CO; NM; NV; TX; UT; Mexico (Chihuahua)
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AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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Discussion | Euphorbia parryi is similar to E. missurica, differing only by the generally narrow involucral gland appendages and prostrate habit in E. parryi as opposed to the conspicuous involucral gland appendages and ascending-erect habit in E. missurica. Euphorbia parryi has sometimes been considered the western race of E. missurica (D. S. Correll and M. C. Johnston 1970). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 281. | FNA vol. 12, p. 322. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Chamaesyce longeramosa, C. parryi, E. longeramosa | E. exstipulata var. lata |
Name authority | Engelmann: Amer. Naturalist 9: 350. (1875) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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