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Parish's sandmat

Habit Herbs or subshrubs, perennial, with thickened and often woody rootstock. Herbs, annual, with taproot.
Stems

prostrate, sometimes forming dense mounds, 10–50 cm, glabrous.

erect, usually branched near base, 6–22 cm, glabrous.

Leaves

opposite;

stipules distinct, subulate-filiform, 0.3–0.9 mm, pilose;

petiole 0.3–1.2 mm, glabrous;

blade usually ovate, rarely oblong, 2–7 × 1–5 mm, base usually asymmetric, rounded to hemicordate, margins entire, apex usually obtuse, rarely acute, surfaces glabrous;

only midvein conspicuous.

petiole absent;

blade linear to slightly lanceolate or linear-oblanceolate, 5–18 × 0.5–2.5 mm, base linear attenuate, margins entire, apex rounded to obtuse or acute, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

obconic to campanulate, 1–1.4 × 0.9–1.3 mm, glabrous except for pilose lobes;

glands 4, pink to maroon, circular, 0.3–0.4 × 0.3–0.4 mm;

appendages absent.

infundibular, 0.8–1.1 × 0.6–0.9 mm, glabrous;

glands 4, crescent-shaped; 0.2–0.4 × 0.5–0.6 mm;

horns divergent, 0.5–0.7 mm.

Staminate flowers

40–50.

5–10.

Pistillate flowers

ovary glabrous;

styles 0.3–0.6 mm, 2-fid 1/2 length.

ovary glabrous;

styles 0.3–0.5 mm, 2-fid.

Capsules

ovoid to oblate-ovoid, 1.6–1.7 × 1.6–1.9 mm, glabrous;

columella 1.2–1.5 mm.

ovoid-globose, 1.8–2.2 × 3–3.2 mm, slightly 3-lobed;

cocci rounded, smooth, glabrous;

columella 1.5–2 mm.

Seeds

whitish to light brown, ovoid, 4-angled in cross section, 1.2–1.4 × 0.6–0.8 mm, rugose or with indistinct, irregular, low transverse ridges.

white to gray, ellipsoid, 1.4–1.7 × 1–1.3 mm, with deep, irregular to rounded, shallow to concave depressions over entire surface;

caruncle reniform-ovate, depressed-conic, 0.5–0.7 × 0.7–1 mm.

Cyathia

solitary at distal nodes;

peduncle 0.1–0.6(–2.2) mm.

peduncle 0–0.5 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched;

pleiochasial bracts similar in shape to but slightly shorter and wider than distal leaves;

dichasial bracts distinct, not imbricate, reniform-ovate to subdeltate-ovate or broadly ovate-lanceolate, base obliquely truncate to rounded, margins entire, apex obtuse to broadly acuminate;

axillary cymose branches 0–3.

Euphorbia parishii

Euphorbia austrotexana

Phenology Flowering and fruiting fall–summer.
Habitat Desert scrub, often with creosote bush, disturbed roadsides, rocky soils.
Elevation -90–600 m. (-300–2000 ft.)
Distribution
from FNA
CA; NV
[BONAP county map]
from FNA
TX
Discussion

Euphorbia parishii is common in the Death Valley region of southern California, where it is often encountered well below sea level. The species is frequently confused with E. micromera and E. polycarpa but differs from the former in being a more robust plant with larger cyathia and from the latter in lacking involucral gland appendages.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Euphorbia austrotexana occurs in stabilized sandy soil in the south Texas plains (M. H. Mayfield 2013). It is similar to E. longicruris but differs from that species in its often narrowly oblanceolate to linear leaves and its white to gray, ellipsoid seeds that are covered with minute, concave depressions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems 10–22 cm; leaf blades linear-oblanceolate, apices rounded; dichasial bracts reniform-ovate to subdeltate-ovate, bases truncate; seeds 1.6–1.7 × 1.1–1.3 mm, surfaces with regular concave depressions; Atascosa, Bexar, Wilson counties.
var. austrotexana
1. Stems 6–13 cm; leaf blades linear to slightly lanceolate, apices acute; dichasial bracts broadly ovate-lanceolate, bases rounded; seeds 1.4–1.5 × 1–1.1 mm, surfaces with irregular, not obviously concave depressions; Jim Hogg, Kenedy counties.
var. carrii
Source FNA vol. 12, p. 280. FNA vol. 12, p. 298.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. austrotexana var. austrotexana, E. austrotexana var. carrii
Synonyms Chamaesyce parishii, E. polycarpa var. parishii
Name authority Greene: Bull. Calif. Acad. Sci. 2: 56. (1886) Mayfield: J. Bot. Res. Inst. Texas 7: 634, figs. 1, 2[row 3, left & center]. (2013)
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