Euphorbia ophthalmica |
Euphorbia velleriflora |
|
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Florida hammock sandmat |
Caliche sandmat |
|
Habit | Herbs, usually annual, rarely short-lived perennial, with slender to slightly thickened taproot. | Herbs, perennial, with strongly thickened and lignified rootstock. |
Stems | usually prostrate, rarely ascending, 6–22 cm, usually both strigillose and hirsute. |
prostrate, usually mat-forming, terete to slightly flattened and winged, to 30 cm, villous to pilose. |
Leaves | opposite; stipules distinct, subulate-filiform, undivided or divided into 2–4 narrowly triangular to linear-subulate segments, no dark, circular glands at base of stipules, 0.9–1.5 mm, pilose or strigillose; petiole 0.3–1.2 mm, glabrescent, strigillose, or sericeous; blade usually ovate or oblong, rarely subrhombic, 4–13 × 3–7 mm, base asymmetric, one side usually angled and other side rounded, margins coarsely serrulate, apex acute, surfaces often with red spot in center, strigillose or sericeous, or adaxial surface glabrescent; 3-veined from base. |
opposite; stipules distinct or connate at base, 2-fid to laciniate into 2–5 linear to subulate divisions, 0.5–1(–2) mm, glabrous or villous; petiole 0.5–2 mm, villous; blade usually ovate to oblong, rarely suborbiculate, 5–13 × 4–8 mm, base asymmetric, hemicordate, margins serrulate, apex obtuse, surfaces villous; 3-veined from base, usually only midvein conspicuous. |
Involucre | obconic, 0.5–0.7 × 0.4–0.6 mm, strigillose; glands 4, yellow green to pink, circular to slightly oblong, 0.1–0.2 × 0.1–0.2 mm; appendages absent or white to pink, forming thin rim around edge of gland or oblong, 0.1–0.2 × 0.1–0.3 mm, distal margin entire or shallowly lobed. |
turbinate, 0.9–1.4 × 0.7–1.1 mm, villous; glands 4, pink, oblong to reniform, 0.1–0.2 × 0.3–0.4 mm; appendages white to pink, flabellate, ovate, or oblong, 0.3–0.5 × 0.4–0.7 mm, distal margin entire, crenulate, or erose, often puberulent-ciliate. |
Staminate flowers | 2–8. |
8–12. |
Pistillate flowers | ovary strigillose, often canescent when young; styles 0.1–0.3 mm, 2-fid 1/2 to nearly entire length. |
ovary villous; styles 0.2–0.4(–0.5) mm, 2-fid 1/2 length. |
Capsules | ovoid, 1–1.2 × 1–1.3 mm, strigillose; columella 0.7–1.1 mm. |
broadly ovoid, 1.5–1.9 × 1.3–1.7 mm, villous (uniformly so, but most pronounced toward base and along keels); columella 1.2–1.4 mm. |
Seeds | orange-brown to pinkish, narrowly ovoid, 4-angled in cross section, 0.7–0.9(–1.1) × 0.5 mm, usually rugulose, with 3–6 faint, low, transverse ridges, rarely almost smooth. |
white to gray or pink, narrowly pyramidal-ovoid, 4-angled in cross section, 1.1–1.3 × 0.4–0.6 mm, rugulose with shallow depressions separated by inconspicuous transverse ridges. |
Cyathia | in dense, terminal, capitate glomerules, with reduced, bractlike leaves subtending cyathia; peduncles 0–0.8 mm. |
solitary at distal nodes or at nodes of leafy, congested, axillary branches; peduncle 1–1.5(–2.5) mm. |
Euphorbia ophthalmica |
Euphorbia velleriflora |
|
Phenology | Flowering and fruiting year-round. | Flowering and fruiting summer. |
Habitat | Hammock forests, disturbed areas in lawns, roadsides. | Disturbed habitats, particularly roadsides, grasslands, live oak-thorn scrub. |
Elevation | 0–200 m. (0–700 ft.) | 50–80 m. (200–300 ft.) |
Distribution |
AR; CA; FL; GA; LA; MO; PA; Mexico; Central America; South America; West Indies [Introduced in Europe] |
TX; Mexico; Central America (Guatemala) [Introduced in North America] |
Discussion | Euphorbia ophthalmica is a weedy species distributed throughout the Neotropics. It is also adventive in the Old World. Whether it is indeed native to the southeastern United States is questionable; it is introduced in Arkansas, California, Missouri, and Pennsylvania and likely occurs also in other states. Although sometimes treated as E. hirta var. procumbens, E. ophthalmica appears sufficiently distinct to justify recognition at the rank of species, differing primarily by its mostly prostrate growth form, smaller leaves, and strictly terminal clusters of cyathia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia velleriflora is native and widespread from northern Mexico to Guatemala. The species is apparently a recent introduction in southern Texas (W. R. Carr and M. H. Mayfield 1993), as the first records are from the 1990s. It is also expected in southern Arizona due to the presence of collections made a few kilometers south of the border in the Mexican state of Sonora. Euphorbia velleriflora is very similar to E. stictospora, and the two are sometimes confused. The two can be readily distinguished on the basis of their styles: unbranched in E. stictospora and 2-fid in E. velleriflora. Also, the involucral gland appendages of E. velleriflora are ciliate with short hairs, whereas those of E. stictospora are glabrous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 280. | FNA vol. 12, p. 292. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce ophthalmica, E. hirta var. procumbens, E. pilulifera var. procumbens | Anisophyllum velleriflorum, Chamaesyce velleriflora |
Name authority | Persoon: Syn. Pl. 2: 13. (1806) | (Klotzsch & Garcke) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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