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Contura Creek sandmat, Contura Creek spurge, valley spurge

Heller's spurge

Habit Herbs, annual, with taproot. Herbs, annual, with taproot.
Stems

prostrate, 10–35 cm, glabrous or pilose.

ascending, branched proximally, 15–30 cm, glabrous.

Leaves

opposite;

stipules distinct, subulate, 0.5–1.6 mm, glabrous or pilose;

petiole 0.3–2 mm, glabrous or pilose;

blade ovate to deltate or falcate, 2.3–13 × 1.5–6 mm, base asymmetric, usually cordate, rarely rounded, margins occasionally reddish, entire, often revolute, apex acute to obtuse, occasionally mucronate, surfaces glabrous or pilose;

midvein conspicuous, lateral veins frequently visible abaxially.

petiole usually 1–3 mm, absent distally;

blade spatulate, 6–15 × 3–5 mm, base broadly attenuate, margins entire, apex usually obtuse to rounded, sometimes retuse, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

campanulate, 1–2.5 × 1.3–2.4 mm, glabrous or pilose;

glands 4, yellow becoming deep red, elliptic or oblong to orbiculate, 0.4–0.6 × 0.5–0.7 mm;

appendages absent or whitish, orbiculate, 0.1–0.2 × 0.3–0.5 mm, distal margin entire.

campanulate, 0.9–1.2 × 0.7–1 mm, glabrous;

glands 4, elliptic, 0.1–0.2 × 0.3–0.4 mm;

horns slightly divergent, 0.1–0.2 mm.

Staminate flowers

30–70.

8–10.

Pistillate flowers

ovary glabrous or pilose;

styles 0.4–0.5 mm, 2-fid at apex.

ovary glabrous;

styles 0.4–0.5 mm, 2-fid.

Capsules

subglobose to broadly ovoid, 1.4–2.7 × 1.9–3.1 mm, glabrous or pilose;

columella 1.4–2 mm.

depressed-globose, 2.1–2.5 × 2.5–3 mm, 3-lobed;

cocci flattened, smooth, glabrous;

columella 1.2–1.7 mm.

Seeds

whitish gray to black, ovoid to oblong, terete to bluntly subangled in cross section, 1.1–1.7 × 0.8–1.3 mm, rugose or smooth.

white to light gray, ovoid, 1.4–1.6 × 1–1.2 mm, smooth;

caruncle 2-lobed, thin, 0.4 × 0.7 mm.

Cyathia

solitary at distal nodes;

peduncle 0.9–2.2 mm.

peduncle 0.2–0.4 mm.

Cyathial

arrangement: terminal pleiochasial branches 3, each 1–5 times 2-branched;

pleiochasial bracts oblong, similar in size to distal leaves;

dichasial bracts distinct, orbiculate-ovate to nearly reniform, subpandurate, base broadly cuneate to truncate, margins entire, apex obtuse, mucronate;

axillary cymose branches 0–5.

Euphorbia ocellata

Euphorbia helleri

Phenology Flowering and fruiting late winter–spring.
Habitat Forests, stream banks, roadsides, shaded areas with sandy, calcareous soils.
Elevation 0–50 m. (0–200 ft.)
Distribution
from FNA
AZ; CA; ID; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; TX; Mexico (Nuevo León)
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The smooth, white to light gray seeds of Euphorbia helleri easily distinguish it from other annual members of subg. Esula in North America. Collections of E. helleri have been made near Brownsville, Texas, and thus it is possible that the species occurs in northern Tamaulipas, Mexico. The Louisiana record (Webster Parish) likely represents introduced plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems, leaves, ovaries, and capsules glabrous; involucral gland appendages absent.
subsp. ocellata
1. Stems, leaves, ovaries, and capsules pilose; involucral gland appendages present.
subsp. rattanii
Source FNA vol. 12, p. 279. FNA vol. 12, p. 303.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. ocellata subsp. ocellata, E. ocellata subsp. rattanii
Synonyms Chamaesyce ocellata Tithymalus helleri
Name authority Durand & Hilgard: Pl. Heermann., 46. (1854) Millspaugh: Bot. Gaz. 26: 268, fig. [p. 270]. (1898)
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