Euphorbia oblongata |
Euphorbia polygonifolia |
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Balkan spurge, egg-leaf spurge, oblong spurge |
dune spurge, euphorbe à feuilles de renouée, seaside sandmat, seaside spurge |
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Habit | Herbs, perennial, with woody taproot. | Herbs, annual, with taproot. |
Stems | erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches). |
usually prostrate, occasionally ascending, 5–30 cm, glabrous. |
Leaves | petiole absent; blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous; venation inconspicuously pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally (distal portion of stem), triangular-subulate, entire or divided, 0.8–1.5 mm, glabrous; petiole 1–3 mm, glabrous; blade oblong, linear-oblong, or linear-lanceolate, 5–16 × 2–4 mm, base slightly asymmetric, obtuse or subcordate, margins entire, apex obtuse, often mucronulate, surfaces uniformly green or reddish tinged, glabrous; venation obscure. |
Involucre | cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous; glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm; horns absent. |
obconic-campanulate, 1.2–1.7 × 1–1.4 mm, glabrous; glands 4, occasionally rudimentary, green-yellow to tan or orange-tinged, occasionally shortly stipitate, usually broadly oval to subcircular, sometimes figure eight-shaped, shallowly cupped, 0.1–0.3 × 0.2–0.4 mm; appendages absent or rudimentary. |
Staminate flowers | 15–40. |
5–14. |
Pistillate flowers | ovary glabrous; styles 1.5–2 mm, 2-fid. |
ovary glabrous; styles 0.7–1 mm, 2-fid 1/2 length. |
Capsules | globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed; cocci rounded, verrucose-tuberculate, glabrous; columella 2.5–3.3 mm. |
broadly ovoid, 3–3.5(–4) × (2–)2.4–3 mm, glabrous; columella 2–3 mm. |
Seeds | brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm. |
ashy white, wedge-shaped to slightly ovoid, weakly dorsiventrally compressed and elliptic-terete to bluntly subangled in cross section, back strongly rounded, face slightly rounded, (2–)2.2–2.8 × 1.6–1.9 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched; pleiochasial bracts ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 1–5 mm. |
solitary or in small, cymose clusters at distal nodes; peduncle 0.5–5 mm. |
Euphorbia oblongata |
Euphorbia polygonifolia |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting early summer–fall. |
Habitat | Waste areas, disturbed sites, roadsides, fields, pastures. | Sandy maritime and freshwater beaches and foredunes. |
Elevation | 30–900 m. (100–3000 ft.) | 0–200 m. (0–700 ft.) |
Distribution |
CA; OR; WA; s Europe [Introduced in North America]
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CT; DE; FL; GA; IL; IN; MA; MD; ME; MI; NC; NH; NJ; NY; OH; PA; RI; SC; VA; WI; NB; NS; ON; PE; QC
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Discussion | Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia polygonifolia is native to coastal beaches and dunes along the Atlantic Ocean from the maritime provinces of Canada south to northern Florida. The species also occurs disjunctly along the shores of the North American Great Lakes. Euphorbia polygonifolia was native to Quebec but is now considered extirpated from the province. It has apparently been introduced in Europe, but it is unclear if it has persisted there (L. C. Wheeler 1941). Immature individuals of this species can be somewhat difficult to distinguish from E. bombensis where their ranges overlap (Virginia to northern Florida). Where they occur together, E. polygonifolia tends to be a pioneer species on the upper beach and foredune front, whereas E. bombensis tends to inhabit areas behind the foredune (R. D. Porcher and D. A. Rayner 2002). Euphorbia polygonifolia can be distinguished also by its larger capsules and larger, wedge-shaped to slightly ovoid seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 284. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus oblongatus | Chamaesyce polygonifolia |
Name authority | Grisebach: Spic. Fl. Rumel. 1: 136. (1843) | Linnaeus: Sp. Pl. 1: 455. (1753) |
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