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Balkan spurge, egg-leaf spurge, oblong spurge

Boquillas sandmat, canyon spurge, swallow spurge

Habit Herbs, perennial, with woody taproot. Herbs, annual, with slender taproot.
Stems

erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches).

prostrate, 5–35 cm, glabrous.

Leaves

petiole absent;

blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous;

venation inconspicuously pinnate, midvein prominent.

opposite;

stipules distinct, subulate, 0.5–0.8 mm, glabrous;

petiole 0.8–1 mm, glabrous;

blade oblong, ovate-oblong, to narrowly elliptic-oblong, 5–11.5 × 1–4 mm, base asymmetric, cuneate to rounded, margins entire, thickened and often revolute on drying, apex obtuse, surfaces glabrous;

only midvein conspicuous.

Involucre

cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous;

glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm;

horns absent.

turbinate, 0.9–1.5 × 0.8–1.3 mm, glabrous;

glands 4, occasionally rudimentary, red to purple, deeply concave, subcircular, 0.3–0.4 × 0.3–0.4 mm;

appendages white, semilunate to slightly flabellate, 0.1–0.3 × 0.5–0.8 mm, distal margin entire.

Staminate flowers

15–40.

28–40.

Pistillate flowers

ovary glabrous;

styles 1.5–2 mm, 2-fid.

ovary glabrous;

styles 0.3–0.4 mm, 2-fid nearly entire length.

Capsules

globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed;

cocci rounded, verrucose-tuberculate, glabrous;

columella 2.5–3.3 mm.

broadly ovoid, 1.7–2 × 1.5–1.6 mm, glabrous;

columella 1.4–1.7 mm.

Seeds

brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm.

narrowly pyramidal-ovoid, 4-angled in cross section, 1.3–1.5 × 0.6–0.7 mm, with very faint transverse ridges or wrinkles.

Cyathial

arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched;

pleiochasial bracts ovate, similar in size to distal leaves;

dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate;

axillary cymose branches 0–4.

Cyathia

peduncle 1–5 mm.

solitary at distal nodes;

peduncle 0.9–1.5 mm.

Euphorbia oblongata

Euphorbia golondrina

Phenology Flowering and fruiting spring–fall. Flowering and fruiting late spring–fall.
Habitat Waste areas, disturbed sites, roadsides, fields, pastures. Deep, sandy riverbanks.
Elevation 30–900 m. (100–3000 ft.) 600 m. (2000 ft.)
Distribution
from FNA
CA; OR; WA; s Europe [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila)
[BONAP county map]
Discussion

Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia golondrina was first collected on a sandy riverbank at the entrance of Boquillas Canyon on the Texas/Mexico border, and the species appears to be restricted to the vicinity of the Rio Grande in western Texas and northern Mexico. Aside from the type locality at Boquillas Canyon, E. golondrina has been documented from additional locations along the Rio Grande in Brewster, Hudspeth, and Presidio counties in Texas and from northern Coahuila and Chihuahua, Mexico. Phylogenetic data place E. golondrina in a clade of primarily Chihuahuan desert annual and perennial herbaceous species (for example, E. chaetocalyx, E. fendleri, E. perennans, E. simulans, E. spurca, and E theriaca; Y. Yang and P. E. Berry 2011). Euphorbia golondrina is superficially similar to other glabrous species in western Texas (for example, E. micromera and E. theriaca), but E. micromera and E. theriaca either lack involucral gland appendages or have shorter, triangular appendages compared to the typical semilunate appendages in E. golondrina.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 305. FNA vol. 12, p. 269.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalus oblongatus Chamaesyce golondrina
Name authority Grisebach: Spic. Fl. Rumel. 1: 136. (1843) L. C. Wheeler: Proc. Biol. Soc. Wash. 53: 8. (1940)
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