Euphorbia oblongata |
Euphorbia cumulicola |
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Balkan spurge, egg-leaf spurge, oblong spurge |
coastal dune sandmat |
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Habit | Herbs, perennial, with woody taproot. | Herbs, usually annual, rarely perennial, with taproot. |
Stems | erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches). |
prostrate, 10–20 cm, glabrous. |
Leaves | petiole absent; blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous; venation inconspicuously pinnate, midvein prominent. |
opposite; stipules distinct, linear-subulate, usually divided into 3–7 linear segments, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade narrowly elliptic to lanceolate, 4–8 × 1–2 mm uniform in size, base asymmetric, obtuse to rounded, margins entire, apex obtuse to acute, surfaces green to reddish flushed, glabrous; only midvein conspicuous. |
Involucre | cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous; glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm; horns absent. |
campanulate, 0.9–1.1 × 1–1.3 mm, glabrous; glands 4, green to red, slightly stipitate, subcircular, 0.1 × 0.1 mm; appendages white or pink, fringing edge of gland, short-flabellate, often rudimentary, 0.1–0.2 × 0.1 mm, distal margin crenate or entire. |
Staminate flowers | 15–40. |
5–8. |
Pistillate flowers | ovary glabrous; styles 1.5–2 mm, 2-fid. |
ovary glabrous; styles 0.2–0.3 mm, 2-fid at apex to nearly 1/2 length. |
Capsules | globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed; cocci rounded, verrucose-tuberculate, glabrous; columella 2.5–3.3 mm. |
ovoid or subglobose, 1.8–2 × 2–2.3 mm, glabrous; columella 1.2–1.5 mm. |
Seeds | brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm. |
white to gray-brown, ovoid, terete to bluntly subangled in cross section, 1–1.3 × 1 mm, smooth or minutely pitted, with smooth brown line from top to bottom on adaxial side. |
Cyathial | arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched; pleiochasial bracts ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate; axillary cymose branches 0–4. |
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Cyathia | peduncle 1–5 mm. |
solitary at distal nodes; peduncle 1 mm. |
Euphorbia oblongata |
Euphorbia cumulicola |
|
Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting year-round. |
Habitat | Waste areas, disturbed sites, roadsides, fields, pastures. | Sandy oak hammocks, open sandy areas behind mangroves, disturbed sandy sites. |
Elevation | 30–900 m. (100–3000 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
CA; OR; WA; s Europe [Introduced in North America]
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FL |
Discussion | Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia cumulicola could be confused with E. bombensis, both of which are widespread in Florida, but there the latter occurs on beaches close to the ocean, whereas E. cumulicola is not a beach-inhabiting species but occurs in more protected sandy habitats such as hammocks or stabilized dunes behind mangroves. Also, E. bombensis has fleshier leaves and larger seeds, and is more compact in habit than E. cumulicola, which is a more sprawling and densely branched plant with leaves that are usually purplish tinged along the margins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 305. | FNA vol. 12, p. 265. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus oblongatus | Chamaesyce cumulicola |
Name authority | Grisebach: Spic. Fl. Rumel. 1: 136. (1843) | (Small) Oudejans: Phytologia 67: 45. (1989) |
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